BSc (Hons), MSc, PhD (Cape Town)

John Day Building: 3.32

Activities and research interests

General ornithology, seabird-fishery interactions, evolutionary ecology (focusing on resolving closely-related species complexes), marine debris, solid waste management, the biology of oceanic islands.

Peter Ryan graduated from the University of Cape Town, researching the impacts of ingested plastic on seabirds (MSc) and the evolutionary ecology of buntings in the Tristan archipelago (PhD). After a post-doc at the University of California where he taught a senior course in ornithology at UC Davis, he returned to UCT in 1993 as Academic Coordinator of the Masters Programme in Conservation Biology at the Fitztitute. Although he mostly works on seabirds and their conservation, he has wide-ranging interests in avian biology, and still maintains an interest in plastic pollution, especially in marine systems. He has served as the President of BirdLife South Africa and is an associate editor of Antarctic Science and Bird Conservation International. He has been an honorary Conservation Officer at Tristan da Cunha since 1989, and is a member of Tristan’s Biodiversity Advisory Group. Peter has spent more than a year on Inaccessible Island, studying the island's endemic buntings as well as conducting surveys of threatened birds and conducting control programmes against alien plants. He is a keen birder and believes in the importance of promoting amateur involvement in ornithology. He has written several books about birds and their identification, and is scientific advisor to African Birdlife, a popular magazine about birds produced by BirdLife South Africa.

Current students

Doctoral

Robin Colyn: Assessing the distribution and population sizes of range restricted larks across an aridity gradient in southern Africa.

Christie Craig: Understanding the state of the Blue Crane population in the Western Cape (Co-supervisor: Tanya Smith, EWT)

Kyle Maclean: Booms, grids and nets: intercepting macroplastic debris in rivers.

Vonica Perold: Temporal and spatial heterogeneity in marine plastic pollution: seabirds, neuston nets and beach litter.

Kim Stevens: The foraging ecology and breeding success of Grey-Headed albatross Thalassarche chrysostoma  on Marion Island.

Marc Travers: Mitigating powerline collisions by breeding seabirds in Hawaii.

Masters

Chukwudi Nwaigwe: The importance of beach-goers as a source of beach litter.

Peer-reviewed publications

Plastic pollution (76 papers)

Clark, B., Carneiro, A., Pearmain, E., Rouyer, M-M., Clay, T. and 199 others. 2023. Global assessment of marine plastic exposure risk for oceanic birds. Nature Communications 

Suaria, G., Cappa, P., Perold, V., Aliani, S. and Ryan, P.G. 2023. Abundance and composition of floating microplastics in the eastern and southern sectors of the Atlantic Ocean. Marine Pollution Bulletin 193: 115109 (doi:10.1016/j.marpolbul.2023.115109)

Weideman, E.A., Perold, V., Donnarumma, V., Suaria, G. and Ryan, P.G. 2023. Proximity to coast and major rivers influence the density of floating microplastics and other litter in east African coastal waters. Marine Pollution Bulletin 188: 114644 (doi:10.1016/j.marpolbul.2023.114644)

van Emmerik, T.H.M., González-Fernández, D., Laufkötter, C., Blettler, M., Lusher, A., Hurley, R. and Ryan, P.G. 2023. Focus on plastics from land to aquatic ecosystems. Environmental Research Letters 18: 040401 (doi:10.1088/1748-9326/acc086)

Honorato-Zimmer, D., Weideman, E.A., Ryan, P.G. and Thiel, M. 2022. Amounts, sources, fates and ecological impacts of marine litter and microplastics in the western Indian Ocean region: a review and recommendations for actions. Oceanography and Marine Biology: an Annual Review 60: 533-590 (doi:10.1201/9781003288602-11)

Ryan, P.G. and Chitaka, T.Y. 2022. Do we need more research on the environmental impacts of plastics? Environmental Research Letters 17: 090201 (doi:10.1088/1748-9326/ac8ab5)

Robuck, A.R., Hudak, C.A., Agvent, L., Emery, G., Ryan, P.G., Perold, V., Powers, K.D., Pedersen, J., Thompson, M.A., Suca, J., Moore, M.J., Harms, C.A., Bugoni, L., Glass, T., Wiley, D.N. and Lohmann, R. 2022. Birds of a feather eat plastic together: High levels of plastic ingestion in Great Shearwater adults and juveniles across their annual migratory cycle. Frontiers in Marine Science 8: 719721 (doi:10.3389/fmars.2021.719721)

Maclean, K., Weideman, E.A., Perold, V. and Ryan, P.G. 2021. Buoyancy affects stranding rate and dispersal distance of floating litter entering the sea from river mouths. Marine Pollution Bulletin 173: 113028 (doi:10.1016/j.marpolbul.2021.113028)

Yamashita, R., Hiki, N., Kashiwada, F., Takada, H., Mizukawa, K., Hardesty, B.D., Roman, L., Hyrenbach, D., Ryan, P.G., Dilley, B.J., Muñoz-Pérez, J.P., Valle, C.A., Pham, C.K., Frias, J., Nishizawa, B., Takahashi, A., Thiebot, J.-B., Will, A., Kokubun, N., Watanabe, Y.Y., Yamamoto, T., Shiomi, K., Shimabukuro, U. and Watanuki, Y. 2021. Plastic additives and legacy persistent organic pollutants in the preen gland oil of seabirds sampled across the globe. Environmental Monitoring and Contaminants Research 1: 97-112 (doi:10.5985/emcr.20210009)

Ryan, P.G., Weideman, E.A., Perold, V., Hofmeyr, G.J.G. and Connan, M.L. 2021. Message in a bottle: assessing the sources and origins of beach litter to tackle marine pollution. Environmental Pollution 288: 117729 (doi:10.1016/j.envpol.2021.117729)

Connan, M., Perold, V., Dilley, B.J., Barbraud, C., Cherel, Y. and Ryan, P.G. 2021. The Indian Ocean ‘garbage patch’: empirical evidence from floating macro-litter. Marine Pollution Bulletin 169: 112559 (doi:10.1016/j.marpolbul.2021.112559). 

Ryan, P.G. and Perold, V. 2021. Limited dispersal of riverine litter onto nearby beaches during rainfall events. Estuarine, Coastal and Shelf Science 251: 107186 (doi:10.1016/j.ecss.2021.107186)

Ohgaki, T., Takada, H., Yoshida, R., Mizukawa, K., Yeo, B.G., Alidoust, M., Hirai, N., Yamashita, R., Tokumaru, T., Watanabe, I., Onwona-Agyeman, S., Gardiner, P., Eriksen, M., Kelly, J.F., Rodríguez-Sierra, C.J., Colasse, L., Baztan, J., Barretto, F.P., Izar, G.M., Abessa, D., Zakaria, M.P., Kwan, C.S. Saha, M., Ryan, P.G., Weerts, S., Ofosu-Anim, J., Sabi, E.B., Akita, L.G., Tait, H., Eriksson, C., and Burton, H. 2021. International pellet watch: Global monitoring of polybrominated diphenyl ethers (PBDEs) in plastic resin pellets. Environmental Monitoring and Contaminants Research 1: 1-16 (doi:10.5985/emcr.20210002)

Ryan, P.G., Weideman, E.A., Perold, V. and Moloney, C.L. 2020. Towards balancing the budget: surface macroplastics dominate the mass of particulate pollution stranded on beaches. Frontiers in Marine Science 7: 575395 (doi:10.3389/fmars.2020.575395).

Ryan, P.G., Maclean, K. and Weideman, E.A. 2020. The impact of the COVID-19 lockdown on urban street litter in South Africa. Environmental Processes 7: 1303-1312. (doi.org/10.1007/s40710-020-00472-1)

Weideman, E.A., Munroe, C., Perold, V., Omardien, A. and Ryan, P.G. 2020. Ingestion of plastic litter by the sandy anemone Bunodactis reynaudi in South Africa. Environmental Pollution 267: 115543. (doi.org/10.1016/j.envpol.2020.115543)

Ryan, P.G. 2020. Land or sea? What bottles tell us about the origins of beach litter in Kenya. Waste Management 116: 49-57.

Weideman, E.A., Perold, V., Omardien, A., Smyth, L.K. and Ryan, P.G. 2020. Quantifying temporal trends in anthropogenic litter in a rocky intertidal habitat. Marine Pollution Bulletin 160: 111543.

Perold, V., Schoombie, S. and Ryan, P.G. 2020. Decadal changes in plastic litter regurgitated by albatrosses and giant petrels at sub-Antarctic Marion Island. Marine Pollution Bulletin 159: 111471.

Ryan, P.G. 2020. Using photographs to record plastic in seabird nests. Marine Pollution Bulletin 156: 111262.

Ryan, P.G. and Schofield, A. 2020. Low densities of macroplastic debris in the Pitcairn Islands Marine Reserve. Marine Pollution Bulletin 157: 111373.

Provencher, J.F., Liboiron, M., Borrelle, S.B., Bond, A.L., Rochman, C., Lavers, J.L., Avery-Gomm, S., Yamashita, R., Ryan, P.G., Lusher, A.L., Hammer, S., Bradshaw, H., Kahn, J. and Mallory, M.L. 2020. A Horizon Scan of research priorities to inform policies aimed at reducing the harm of plastic pollution to biota. Science of the Total Environment (doi.org/10.1016/j.scitotenv.2020.1393812020)

Suaria, G., Achtypi, A., Perold, V., Lee, J.R., Pierucci, A., Bornman, T.G., Aliani, S. and Ryan, P.G. 2020. A global characterization of microfibers in oceanic surface waters. Science Advances 6: eaay8493. (doi.org/10.1126/sciadv.aay8493)

Weideman, E.A., Perold, V. and Ryan, P.G. 2020. Limited long-distance transport of plastic pollution by the Orange-Vaal River system, South Africa. Science of the Total Environment 727: 138653. (doi.org/10.1016/j.scitotenv.2020.138653)

Weideman, E.A., Perold, V., Arnold, G. and Ryan, P.G. 2020. Quantifying changes in litter loads in urban stormwater run-off from Cape Town, South Africa, over the last two decades. Science of the Total Environment 724: 138310. (doi:10.1016/j.scitotenv.2020.138310)

Tavares, D.C., Moura, J., Acevedo-Trejos, E., Crawford, R., Makhado, A., Lavers, J., Witteveen, M., Ryan, P.G. and Merico, A. 2020. Confidence intervals and sample size for estimating the prevalence of plastic debris in seabird nests. Environmental Pollution 263: 114394. (doi.org/10.1016/j.envpol.2020.114394)

Ryan, P.G. 2020. The transport and fate of marine plastics in South Africa and adjacent oceans. South African Journal of Science 116: 7677. (doi.org/10.17159/sajs.2020/7677)

Ryan, P.G., Pichegru, L., Perold, V. and Moloney, C.L. 2020. Monitoring marine plastics – will we know if we’re making a difference? South African Journal of Science 116: 7678. (doi.org/10.17159/sajs.2020/7678)

van Sebille, E., Aliani, S., Law, K.L., Maximenko, N., Alsina, J., Bagaev, A., Bergmann, M., Chapron, B., Chubarenko, I., Cózar, A., Delandmeter, P.; Egger, M., Fox-Kemper, B., Garaba, S., Goddijn-Murphy, L., Hardesty, D., Hoffman, M., Isobe, A., Jongedijk, C., Kaandorp, M., Khatmullina, L., Koelmans, A., Kukulka, T., Laufkötter, C., Lebreton, L., Lobelle, D., Maes, C., Martinez-Vicente, V., Morales, M., Miguel, A., Poulain-Zarcos, M., Rodriguez, E., Ryan, P.G., Shim, W.J., Suaria, G., Thiel, M., van den Bremer, T. and Wichmann, D. 2020. The physical oceanography of the transport of floating marine debris. Environmental Research Letters 15: 023003. (doi:10.1088/1748-9326/ab6d7d)

Suaria, G., Lee, J.R., Le Bouard, F., Aliani, S. and Ryan, P.G. 2020. Floating macro- and microplastics around the Southern Ocean: results from the Antarctic Circumnavigation Expedition. Environment International 136: 105494. (doi:10.1016/j.envint.2020.105494)

Le Guen, C., Suaria, G., Sherley, R., Ryan, P.G., Aliani, S., Boehme, L. and Brierley, A. 2020. Microplastic study reveals the presence of natural and synthetic fibres in the diet of King Penguins (Aptenodytes patagonicus) foraging from South Georgia. Environment International 134: 105303. (doi:10.1016/j.envint.2019.105303)

Ryan, P.G., Weideman, E.A., Perold, V., Durholz, D. and Fairweather, T.P. 2020. A trawl survey of seabed macrolitter on the South African continental shelf. Marine Pollution Bulletin 150: 110741. (doi:10.1016/j.marpolbul.2019.110741)

Ryan, P.G., Suaria, G., Perold, V., Pierucci, A., Bornman, T.G. and Aliani, S. 2020. Sampling microfibres at the sea surface: the effects of mesh size, sample volume and water depth. Environmental Pollution 258: 113413. (doi:10.1016/j.envpol.2019.113413)

Ryan, P.G., Dilley, B.J., Ronconi, R.A. and Connan, M. 2019. Rapid increase in Asian bottles in the South Atlantic Ocean indicates major debris inputs from ships. Proceedings of the National Academy of Sciences U.S.A. 116: 20892–20897. (doi:10.1073/pnas.1909816116)

Weideman, E.A., Perold, V. and Ryan, P.G. 2019. Little evidence that dams in the Orange–Vaal River system trap micro- and meso-plastics. Marine Pollution Bulletin 149: 110664.

Pfaff, M.C., Logston, R.C., Raemaekers, S.J.P.N., Hermes, J.C, Blamey, L.K., Cawthra, H.C., Colenbrander, D.R., Crawford, R.J.M., Day, E., du Plessis, N., Elwen, S.H., Fawcett, S.E., Jury, M.R., Karenyi, N., Kerwath, S.E., Kock, A.A., Krug, M., Lamberth, S.J., Omardien, A., Pitcher, G.C., Rautenbach, C., Robinson, T.B., Rouault, M., Ryan, P.G., Shillington, F.A., Sowman, M., Sparks, C.C., Turpie, J.K., van Niekerk, L., Waldron, H.N., Yeld, E.M. and Kirkman, S.P. 2019. A synthesis of three decades of socio-ecological change in False Bay, South Africa: setting the scene for multidisciplinary research and management. Elementa: Science of the Anthropocene 7: 32. (doi:10.1525/elementa.367)

Ryan, P.G. 2018. Entanglement of birds in plastics and other synthetic materials. Marine Pollution Bulletin 135: 159-164.

Ryan, P.G., Perold, V., Osborne, A. and Moloney, C.L. 2018. Consistent patterns of debris on South African beaches indicate that industrial pellets and other mesoplastic items mostly derive from local sources. Environmental Pollution 238: 1008-1016.

Reynolds, C. and Ryan, P.G. 2018. Micro-plastic ingestion by waterbirds from contaminated wetlands in South Africa. Marine Pollution Bulletin 126: 330-333.

Dell’Ariccia, G., Phillips, R.A., van Franeker, J.A., Gaidet, N., Catry, P., Granadeiro, J.P., Ryan, P.G. and Bonadonna, F. 2017. Does DMS build-up really explain plastic ingestion by seabirds? A comment on Savoca et al. (2016). Scientific Reports 3(6), e1700526.

Witteveen, M., Brown, M. and Ryan, P.G. 2017. Anthropogenic debris in the nests of Kelp Gulls in South Africa. Marine Pollution Bulletin 114: 699-704.

Ryan, P.G. 2016. Ingestion of plastics by marine organisms. In: Takada, H. and Karapanagioti, H.K. (eds) Hazardous Chemicals Associated with Plastics in the Environment. Handbook of Environmental Chemistry 78: 235-266. (doi:10.1007/698_2016_21)

Fazey, F.M.C. and Ryan, P.G. 2016. Debris size and buoyancy influence the dispersal distance of stranded litter. Marine Pollution Bulletin 110: 371-377.

Ryan, P.G., Cole, G., Spiby, K., Nel, R., Osborne, A. and Perold, V. 2016. Impacts of plastic ingestion on post-hatchling loggerhead turtles off South Africa. Marine Pollution Bulletin 107: 155-160.

Ryan, P.G., de Bruyn, P.J.N. and Bester, M.N. 2016. Regional differences in plastic ingestion among Southern Ocean fur seals and albatrosses. Marine Pollution Bulletin 104: 207-210.

Fazey, F.M.C. and Ryan, P.G. 2016. Biofouling on buoyant marine plastics: an experimental study into the effect of size on surface longevity. Environmental Pollution 210: 354-360.

Ryan, P.G. 2015. Does size and buoyancy affect the long-distance transport of floating debris? Environmental Research Letters 10: 084019.

Ryan, P.G. 2015. How quickly do albatrosses and petrels digest plastic particles? Environmental Pollution 207: 438-440.

Ryan, P.G. 2014. Litter survey detects the South Atlantic ‘garbage patch’. Marine Pollution Bulletin 79: 220-224.

Ryan, P.G., Lamprecht, A., Swanepoel, D. and Moloney, C.L. 2014. The effect of fine-scale sampling frequency on estimates of beach litter accumulation. Marine Pollution Bulletin 88: 249-254.

Ryan, P.G., Musker, S. and Rink, A. 2014. Low densities of drifting litter in the African sector of the Southern Ocean. Marine Pollution Bulletin 89: 16-19.

Eriksen, M., Lebreton, L.C.M., Carson, H.S., Thiel, M., Moore, C.J., Borerro, J.C., Galgani, F., Ryan, P.G. and Reissner, J. 2014. Plastic pollution in the world’s oceans: more than 5 trillion plastic pieces weighing over 250,000 tons afloat at sea. PLoS One 9(12): e111913.

Vegter, A.C., Barletta, M., Beck, C., Borrero, J., Burton, H., Campbell, M.L., Eriksen, M., Eriksson, C., Estrades, A., Gilardi, K.V.K., Hardesty, B.D., Ivar do Sul, J.A., Lavers, J.L., Lazar, B., Lebreton, L., Nichols, W.J., Ribic, C.A., Ryan, P.G., Schuyler, Q.A., Smith, S.D.A., Takada, H., Townsend, K.A., Wabnitz, C.C.C., Young, L.C. and Hamann, M. 2014. Global research priorities to mitigate plastic pollution impacts on marine wildlife. Endangered Species Research 25: 225-247.

Ryan, P.G. 2013. A simple technique for counting marine debris at sea reveals steep litter gradients between the Straits of Malacca and the Bay of Bengal. Marine Pollution Bulletin 69: 128-136.

Ryan, P.G., Bouwman, H., Moloney, C.L., Yuyama, M. and Takada, H. 2012. Long-term decreases in persistent organic pollutants in South African coastal waters detected from beached polyethylene pellets. Marine Pollution Bulletin 64: 2756-2760.

Ryan, P.G., Moore, C.J., van Franeker, J.A. and Moloney, C.L. 2009. Monitoring the abundance of plastic debris in the marine environment. Philosophical Transactions of the Royal Society B 364: 1999-2012.

Ryan, P.G. 2008. Seabirds indicate decreases in plastic pellet litter in the Atlantic and south-western Indian Ocean. Marine Pollution Bulletin 56: 1406-1409.

Cliff, G., Dudley, S.F.J., Ryan, P.G. and Singleton, N. 2002. Large sharks and plastic debris in KwaZulu-Natal, South Africa. Marine and Freshwater Research 53: 575-581.

Ballance, A., Ryan, P.G. and Turpie, J.K. 2000. How much is a clean beach worth? The impact of litter on beach users in the Cape Peninsula, South Africa. South African Journal of Science 96: 210-213.

Ryan, P.G. 1996. Plastic litter in marine systems: impacts, sources and solutions. Plastics Southern Africa 26 (6): 20-28.

Ryan, P.G. 1996. Policy recommendations for controlling plastic litter. Plastics Southern Africa 25 (10): 46.

Ryan, P.G. and Swanepoel, D. 1996. Cleaning beaches: sweeping litter under the carpet. South African Journal of Science 92: 275-276.

Ryan, P.G., Swanepoel, D., Rice, N. and Preston, G. 1996. The ‘free’ shopping bag debate: costs and attitudes. South African Journal of Science 92: 163-165.

Ryan, P.G. and Moloney, C.L. 1993. Marine litter keeps increasing. Nature 361: 23.

Ryan, P.G. and Moloney, C.L. 1990. Plastic and other artefacts on South African beaches: temporal trends in abundance and composition. South African Journal of Science 86: 450-452.

Ryan, P.G. 1988. Intraspecific variation in plastic ingestion by seabirds and the flux of plastic through seabird populations. Condor 90: 446-452.

Ryan, P.G. 1988. The characteristics and distribution of plastic particles at the sea-surface off the southwestern Cape Province, South Africa. Marine Environmental Research 25: 249-273.

Ryan, P.G. 1988. Effects of plastic ingestion on seabird feeding: evidence from chickens. Marine Pollution Bulletin 19: 125-128.

Ryan, P.G., Connell, A.D. and Gardner, B.D. 1988. Plastic ingestion and PCBs in seabirds: is there a relationship? Marine Pollution Bulletin 19: 174-176.

Ryan, P.G. and Fraser, M.W. 1988. The use of Great Skua pellets as indicators of plastic pollution in seabirds. Emu 88: 16-19.

Ryan, P.G. and Watkins, B.P. 1988. Accumulation of stranded plastic objects and other artefacts at Inaccessible Island, central South Atlantic Ocean. South African Journal of Antarctic Research 18: 11-13.

Ryan, P.G. 1987. The incidence and characteristics of plastic particles ingested by seabirds. Marine Environmental Research 23: 175-206.

Ryan, P.G. 1987. The origin and fate of artefacts stranded on islands in the African sector of the Southern Ocean. Environmental Conservation 14: 341-346.

Ryan, P.G. 1987. The effects of ingested plastic on seabirds: correlations between plastic load and body condition. Environmental Pollution 46: 119-125.

Ryan, P.G. and Jackson, S. 1987. The lifespan of ingested plastic particles in seabirds and their effect on digestive efficiency. Marine Pollution Bulletin 18: 217-219.

Ryan, P.G. 1986. Seabirds as water quality monitors. Marine Pollution Bulletin 17: 435-436.

Seabird monitoring and conservation (58 papers)

Green, C-P., Green, D.B., Ratcliffe, N., Thompson, D., Lea, M-A., Baylis, A.M.M., Bond, A.L., Bost, C-A., Crofts, S., Cuthbert, R.J., González-Solís, J., Morrison, K.W., Poisbleau, M., Pütz, K., Raya Rey, A., Ryan, P.G., Sagar, P.M., Steinfurth, A., Thiebot, J-B., Tierney, M., Whitehead, T.O., Wotherspoon, S. and Hindell, M.A. 2022. Potential for redistribution of post-moult habitat for Eudyptes penguins in the Southern Ocean under future climate conditions. Global Change Biology 29: 648-667 (doi:10.1111/gcb.16500).

Connan, M., Schoombie, S., Schoombie, J., Dilley, B. and Ryan, P.G. 2022. Natural recolonization of sub-Antarctic Marion Island  by Common Diving Petrels. Ostrich 93: 271-279 (doi:10.2989/00306525.2022.2150706).

Oppel, S., Clark, B., Risi, M., Horswill, C., Converse, S., Jones, C., Osborne, A., Stevens, K., Perold, V., Bond, A., Wanless, R., Cuthbert, R., Cooper, J. and Ryan, P.G. 2022. Cryptic population decrease due to invasive species predation in a long‐lived seabird supports need for eradication. Journal of Applied Ecology 59: 2059-2070 (doi:10.1111/1365-2664.14218).

Ryan, P.G. and Oppel, S. 2022. Winter colony attendance by adult Southern Giant Petrels: implications for rodent eradications. Marine Ornithology 50: 1-4.

Reissinger, R.R., Corney, S., Raymond, B., Lombard, A.T., Bester, M.N., Crawford, R.J.M., Davies, D., de Bruyn, P.J.N., Dilley, B.J., Kirkman, S.P., Makhado, A.B., Ryan, P.G., Schoombie, S., Stevens, K.L., Tosh, C.A., Wege, M., Whitehead, T.O., Sumner, M.D., Wotherspoon, S., Friedlaender, A.S., Cotté, C., Hindell, M.A., Ropert-Coudert, Y., Pistorius, P.A. 2022. Habitat model forecasts suggest potential redistribution of marine predators in the southern Indian Ocean. Diversity and Distributions 28: 142-159 (doi:10.1111/ddi.13447)

Ryan, P.G., Ryan, E.M. and Glass, J.P. 2021. Dazzled by the light: the impact of light pollution from ships on seabirds at Tristan da Cunha. Ostrich 92: 218-224 (doi:10.2989/00306525.2021.1984998)

Wakefield, E.D., Miller, D.L., Bond, S.L., le Bouard, F., Carvalho, P.C., Catry, P., Dilley, B.J., Fifield, D.A., Gjerdrum, C., González-Solís, J., Hogan, H., Laptikhovsky, V., Merkel, B., Miller, J.A.O., Miller, P.I., Pinder, S.J., Pipa, T., Ryan, P.G., Thompson, L.A., Thompson, P.M. and Matthiopoulos, J. 2021. The summer distribution, habitat associations and abundance of seabirds in the sub-polar frontal zone of the Northwest Atlantic. Progress in Oceanography 198. (doi:10.1016/j.pocean.2021.102657)

Davies, T., Carneiro, A.P.B., Tarzia, M., Wakefield, E., Hennicke, J., Frederiksen, M., Hansen, E.S., Campos, B., Hazin, C., Lascelles, B., Anker-Nilssen, T., Arnardóttir, H., Barrett, R.T., Biscoito, M., Bollache, L., Boulinier, T., Catry, P., Ceia, F.R., Chastel, O., Christensen-Dalsgaard, S., Cruz-Flores, M., Danielsen, J., Daunt, F., Dunn, E., Egevang, C., Fagundes, A.I., Fayet, A.L., Fort, J., Furness, R.W., Gilg, O., González-Solís, J., Granadeiro, J.P., Grémillet, D., Guilford, T., Hanssen, S.A., Harris, M.P., Hedd, A., Huffeldt, N.P., Jessopp, M., Kolbeinsson, Y., Krietsch, J., Lang, J., Linnebjerg, J.F., Lorentsen, S-H., Madeiros, J., Magnusdottir, E., Mallory, M.L., Tranquilla, L.M., Merkel, F.R., Militão, T., Moe, B., Montevecchi, W.A., Morera-Pujol, V., Mosbech, A., Neves, V., Newell, M.A., Olsen, B., Paiva, V.H., Peter, H-U., Petersen, A., Phillips, R.A., Ramírez, I., Ramos, J.A., Ramos, R., Ronconi, R.A., Ryan, P.G., Schmidt, N.M., Sigurðsson, I.A., Sittler, B., Steen, H., Stenhouse, I.J., Strøm, H., Systad, G.H.R., Thompson, P., Thórarinsson, T.L., van Bemmelen, R.S.A., Wanless, S., Zino, F. and Dias, M.P. 2021. Multi-species tracking reveals a major seabird hotspot in the North Atlantic. Conservation Letters 2021: e12824. (doi:10.1111/conl.12824)

Beal, M., Dias, M.P., Phillips, R.A., Oppel, S., Hazin, C., Pearmain, E.J., Adams, J., Anderson, D.J., Antolos, M., Arata, J.A., Arcos, J.M., Arnould, J.P.Y., Awkerman, J., Bell, E., Bell, M., Carey, M., Carle, R., Clay, T.A., Cleeland, J., Colodro, V., Connors, M., Cruz-Flores, M., Cuthbert, R., Delord, K., Deppe, L., Dilley, B.J., Dinis, H., Elliott, G., De Felipe, F., Felis, J., Forero, M.G., Freeman, A., Fukuda, A., González-Solís, J., Granadeiro, J.P., Hedd, A., Hodum, P., Igual, J.M., Jaeger, A., Landers, T.J., Le Corre, M., Makhado, A., Metzger, B., Militao, T., Montevecchi, W.A., Morera-Pujol, V., Navarro-Herrero, L., Nel, D., Nicholls, D., Oro, D., Ouni, R., Ozaki, K., Quintana, F., Ramos, R., Reid, T., Reyes-González, M., Robertson, C., Robertson, G., Romdhane, S., Ryan, P.G., Sagar, P., Sato, F., Schoombie, S., Scofield, R.P., Shaffer, S.A., Shah, N.J., Stevens, K.L., Surman, C., Suryan, R.M., Takahashi, A., Tatayah, V., Taylor, G., Thompson, D.R., Torres, L., Walker, K., Wanless, R., Waugh, S.M., Weimerskirch, H., Yamamoto, T., Zajkova, Z., Zango, L. and Carty, P.. 2021. Global political responsibility for the conservation of albatrosses and large petrels. Science Advances 7: eabd7225. (doi:10.1126/sciadv.abd7225)

Dilley, B.J., Hedding, D.W., Schoombie, S. and Ryan, P.G. 2020. A survey of Great-winged Petrels Pterodroma macroptera breeding at sub-Antarctic Marion Island and a revised global population estimate. Ostrich 91: 101-106. (doi:10.2989/00306525.2020.1721587)

Ryan, P.G., Dilley, B.J. and Ronconi, R.A. 2019. Population trends of Spectacled Petrels Procellaria conspicillata and other seabirds at Inaccessible Island. Marine Ornithology 47: 257-265.

Dilley, B.J., Davies, D., Mitham, A., Glass, T., Repetto, J., Swain, G. and Ryan, P.G. 2019. Population estimates of burrow-nesting petrels breeding at the Nightingale Island Group, Tristan da Cunha Archipelago. Marine Ornithology 47: 267-275.

Sherley, R.B., Crawford, R.J.M., Dyer, B.M., Kemper, J., Makhado, A.B., Masotla, M., Pichegru, L., Pistorius, P.A., Roux, J.-P., Ryan, P.G., Tom, D., Upfold, L. and Winker, H. 2019. The status and conservation of Cape Gannets Morus capensis. Ostrich 90: 335-346. (doi.org/10.2989/00306525.2019.1684396)

Dilley, B.J., Hedding, D.W., Henry, D.A.W., Rexer-Huber, K, Parker, G.C., Schoombie, S., Osborne, A. and Ryan, P.G. 2019. Clustered or dispersed: testing the effect of sampling strategy to census burrow-nesting petrels with varied distributions at sub-Antarctic Marion Island. Antarctic Science 31: 231-242. (doi.org/10.1017/S0954102019000300)

Ropert-Coudert, Y., Chiaradia, A., Ainley, D., Barbosa, A., Boersma, P.D., Brasso, R., Dewar, M., Ellenberg, U., Borboroglu, P.G., Emmerson, L., Hickcox, R., Jenouvrier, S., Kato, A., McIntosh, R.R., Lewis, P., Ramirez, F., Ruoppolo, V., Ryan, P.G., Seddon, P.J., Sherley, R.B., Vanstreels, R.E.T., Waller, L., Woehler, E.J. and Trathan, P.N. 2019. Happy feet in a hostile world? The future of penguins depends on proactive management of current and predictable threats. Frontiers in Marine Science 6: 248. (doi:10.3389/fmars.2019.00248)

Rodríguez, A., Arcos, J.M., Bretagnolle, V., Dias, M.P., Holmes, N.D., Louzao, M., Provencher, J., Raine, A.F., Ramírez, F., Rodríguez, B., Ronconi, R.A., Taylor, R.S., Bonnaud, E., Borrelle, S., Cortés, V., Descamps, S., Friesen, V.L., Genovart, M., Hedd, A., Hodum, P., Humphries, G., Le Corre, M., Lebarbenchon, C., Martin, R., Melvin, E.F., Montevecchi, W.A., Pinet, P., Pollet, I.L., Ramos, R., Russell, J.C., Ryan, P.G., Sanz-Aguilar, A., Spatz, D., Travers, M., Votier, S., Wanless, R., Woehler, E. and Chiaradia, A. 2019. Future directions in conservation research on petrels and shearwaters. Frontiers in Marine Science 6: 94 (doi:10.3389/fmars.2019.00094)

Weimerskirch, H., Le Bouard, F., Ryan, P.G. and Bost, C.-A. 2018. Massive decline of one of the world’s largest penguin colonies: the King Penguin colony at Ile aux Cochons, Crozet. Antarctic Science 30: 236-242.

Weimerskirch, H., Delord, K., Fretwell, P.T., Marteau, C., Ryan, P.G. and Barbraud, C. 2018. Status and trends of albatrosses in the French Southern Territories, western Indian Ocean. Polar Biology 41: 1963-1972.

Dias, M., Oppel, S., Bond, A.L., Carneiro, A.P.B., Cuthbert, R.J., Gonzáles-Solís, J., Wanless, R.M., Glass, T., Lascelles, B., Small, C., Phillips, R.A. and Ryan, P.G. 2017. Using globally threatened pelagic birds to identify priority sites for marine conservation in the South Atlantic Ocean. Biological Conservation 211: 76-84.

Krüger, L., Ramos, J.A., Xavier, J.C., Grémillet, D., González-Solís, J., Kolbeinsson, Y., Militãò, T., Navarro, J., Petry, M.V., Phillips, R.Z., Ramírez, I., Reyes-González, J.M., Ryan, P.G., Siguròsson, I.A., van Sebille, E., Wanless, R.M. and Paiva, V.H. 2017. Identification of candidate pelagic marine protected areas through a seabird seasonal-, multispecific- and extinction risk-based approach. Animal Conservation 20: 409-424.

Dilley, B.J., Davies, D., Connan, M., Schramm, M. and Ryan, P.G. 2017. The distribution and abundance of Blue Petrels (Halobaena caerulea) breeding at Subantarctic Marion Island. Emu 117: 222-232.

Schoombie, S., Crawford, R.J.M., Makhado, A.B., Dyer, B.M. and Ryan, P.G. 2016. Recent population trends of Sooty and Light-mantled Albatrosses breeding on Marion Island. African Journal of Marine Science 38: 119-127.

Whittington, P.A., Crawford, R.J.M., Martin, A.P., Randall, R.M., Brown, M., Makhado, A.B., Dyer, B.M., Upfold, L., Harrison, K.H., Huisamen, J., Ryan, P.G., Waller, L. and Witteveen, M. 2016. Recent trends of the Kelp Gull Larus dominicanus in South Africa. Waterbirds 39(sp1): 99-113.

Pichegru, L., Edwards, T.B., Dilley, B.J., Flower, T.P. and Ryan, P.G. 2016. African Penguin tolerance to humans depends on historical exposure at colony level. Bird Conservation International 26: 307-322.

Ryan, P.G., Dilley, B.J., Jones, C. and Bond, A. 2015. Blue Petrels breeding on Gough Island. Ostrich 89: 193-194.

Cuthbert, R.J., Cooper, J. and Ryan, P.G. 2014. Population trends and breeding success of albatrosses and giant petrels at Gough Island in the face of at-sea and on-land threats. Antarctic Science 26: 163-171.

Rexer-Huber, K., Parker, G.C., Ryan, P.G. and Cuthbert, R.J. 2014. Burrow occupancy and population size in the Atlantic Petrel: a comparison of methods. Marine Ornithology 42: 137-141.

Crawford, R.J.M., Randall, R.M., Whittington, P.A., Waller, L., Dyer, B.M., Allan, D.G., Fox, C., Martin, A.P., Upfold, L., Visagie, J., Bachoo, S., Bowker, M., Downs, C.T., Fox, R., Huisamen, J., Makhado, A.B., Oosthuizen, W.H., Ryan, P.G., Taylor, R.H. and Turpie, J.K. 2013. South Africa's coastal-breeding White-breasted Cormorants: population trends, breeding season and movements, and diet. African Journal of Marine Science 35: 473-490.

Ryan, P.G. and Kerr, J. 2012. Is fur seal predation driving the decrease in Northern Rockhopper Penguins Eudyptes moseleyi at Gough Island? Marine Ornithology 40: 73-75.

Ryan, P.G., Dilley, B.J. and Jones, M.G.W. 2012. The distribution and abundance of White-chinned Petrels (Procellaria aequinoctialis) breeding at the sub-Antarctic Prince Edward Islands. Polar Biology 35: 1851-1859.

Lewis, S.F.F., Turpie, J.K. and Ryan, P.G. 2012. Are African Penguins worth saving? The ecotourism value of the Boulders Beach colony. African Journal of Marine Science 34: 497-504.

Lewison, R., Oro, D., Godley, B., Underhill, L., Bearhop, S., Wilson, R., Ainley, D., Arcos, J.M., Boersma, P.D., Borboroglu, P., Boulinier, T., Frederiksen, M., Genovart, M., González-Solís, J., Green, J.A., Grémillet, D., Hamer, K.C., Hilton, G.M., Hyrenbach, K.D., Martínez-Abraín, A., Montevecchi, W.A., Phillips, R.A., Ryan, P.G., Sagar, P., Sydeman, W.J., Yorio, P., Wanless, S., Watanuki, Y. and Weimerskirch, H. 2012. Research priorities for seabirds: improving seabird conservation and management in the 21st century. Endangered Species Research 17: 93-121.

Ryan, P.G. and Ronconi, R.A. 2011. Continued increase in numbers of Spectacled Petrels Procellaria conspicillata. Antarctic Science 23: 332-336.

Crawford, R.J.M., Altwegg, R., Barham, B.J., Barham, P.J., Durant, J.M., Dyer, B.M., Geldenhuys, D., Makhado, A.B., Pichegru, L., Ryan, P.G., Underhill, L.G., Upfold, L., Visagie, J., Waller, L.J. and Whittington, P.A. 2011. Collapse of South Africa’s penguins in the early 21st century. African Journal of Marine Science 33: 139-156.

Robson, B., Glass, T., Glass, N., Glass, J., Green, J., Repetto, C., Rodgers, G., Ronconi, R.A., Ryan, P.G., Swain, G. and Cuthbert, R.J. 2011. Revised population estimate and trends for the endangered Northern Rockhopper Penguin Eudyptes moseleyi at Tristan da Cunha. Bird Conservation International 21: 454-459.

Mwema, M.M., de Ponte Machado, M. and Ryan, P.G. 2010. Breeding seabirds at Dassen Island, South Africa: chances of surviving Great White Pelican predation. Endangered Species Research 9: 125-131.

Pichegru, L., Ryan, P.G., Crawford, R.J.M., van der Lingen, C. and Grémillet, D. 2010.  Behavioural inertia places a top marine predator at risk from environmental change in the Benguela upwelling system. Marine Biology 157: 537-544.

Ryan, P.G., Jones, M.G.W., Dyer, B.M., Upfold, L. and Crawford, R.J.M. 2009. Recent population estimates and trends in numbers of albatrosses and giant petrels breeding at the sub-Antarctic Prince Edward Islands. African Journal of Marine Science 31: 409-417.

Ryan, P.G., Whittington, P.A. and Crawford, R.J.M. 2009. A tale of two islands: contrasting fortunes for Subantarctic Skuas at the Prince Edward Islands. African Journal of Marine Science 31: 431-437.

Converse, S.J., Kendall, W.L., Doherty, P.F.Jr. and Ryan, P.G. 2009. Demography of Marion Island Grey-headed Albatross: analysis utilizing advances in mark-recapture models. Auk 126: 77-88.

Crawford, R.J.M., Ryan, P.G., Dyer, B.M. and Upfold, L. 2009. Recent trends in numbers of Crozet Shags breeding at the Prince Edward Islands. African Journal of Marine Science 31: 427-430.

Crawford, R.J.M., Whittington, P.A., Upfold, L., Ryan, P.G., Petersen, S.L., Dyer, B.M. and Cooper, J. 2009. Recent trends in numbers of four species of penguins at the Prince Edward Islands. African Journal of Marine Science 31: 419-426.

Cuthbert, R., Cooper, J., Burle, M.-H., Glass, C.J., Glass, J.P., Glass, T., Hilton, G.M., Sommer, E., Wanless, R.M. and Ryan, P.G. 2009. Population trends and conservation status of the Northern Rockhopper Penguin Eudyptes moseleyi at Tristan da Cunha and Gough Island. Bird Conservation International 19: 109-120.

Hampton, S., Ryan, P.G. and Underhill, L.G. 2009. The effect of flipper banding on the breeding success of African Penguins Spheniscus demersus at Boulders Beach, South Africa. Ostrich 80: 77-80.

Whittington, P.A., Crawford, R.J.M., Dyer, B.M. and Ryan, P.G. 2009. Estimates of numbers of kelp gulls and Kerguelen and Antarctic terns breeding at the Prince Edward Islands, 1996/97–2008/09. African Journal of Marine Science 31: 439-444.

Ryan, P.G., Dorse, C. and Hilton, G.M. 2006. The conservation status of the Spectacled Petrel Procellaria conspicillata. Biological Conservation 131: 575-583.

Lewis, S., Daunt, F., Grémillet, D., Ryan, P.G., Crawford, R.J.M. and Wanless, S. 2006. Using behavioural variables to identify proximate causes of population change in a seabird. Oecologia 147: 606-614.

Cuthbert, R.J., Sommer, E.S., Ryan, P.G., Cooper, J., and Hilton, G. 2004. Demography and conservation status of the Tristan Albatross Diomedea [exulans] dabbenena. Biological Conservation 117: 471-481.

Ryan, P.G., Cooper, J., Dyer, B.M., Underhill, L.G., Crawford, R.J.M. and Bester, M.N. 2003. Counts of surface-nesting seabirds breeding at sub-Antarctic Prince Edward Island, summer 2001/02. African Journal of Marine Science 25:  441-451.

Crawford, R.J.M., Cooper, J., Dyer, B.M., Greyling, M.D., Klages, N.T.W., Ryan, P.G., Petersen, S.L., Underhill, L.G., Upfold, L., Wilkinson, W., de Villiers, M.S., du Plessis, S., du Toit, M., Leshoro, T.M., Makhado, A.B., Mason, M., Merkle, D., Tshingana, D., Ward, V.L. and Whittington, P.A. 2003. Populations of surface-nesting seabirds at Marion Island, 1994/95-2002/03. African Journal of Marine Science 25: 427-439.

Cuthbert, R.J., Ryan, P.G., Cooper, J. and Hilton, G. 2003. Demography and population trends of the Atlantic Yellow-nosed Albatross Thalassarche chlororhynchos. Condor 105: 439-452.

Ryan, P.G. and Moloney, C.L. 2000. The status of Spectacled Petrels Procellaria conspicillata and other seabirds at Inaccessible Island. Marine Ornithology 28: 93-100.

Ryan, P.G., Avery, G. and Steele, W.K. 1991. Distribution, population sizes and conservation of marine and coastal birds in False Bay. Transactions of the Royal Society of South Africa 47: 649-662.

Ryan, P.G., Dean, W.R.J., Moloney, C.L., Watkins, B.P. and Milton, S.J. 1990. New information on seabirds at Inaccessible Island and other islands in the Tristan da Cunha group. Marine Ornithology 18: 43-54.

Ryan, P.G. and Watkins, B.P. 1988. Birds of the inland mountains of western Dronning Maud Land, Antarctica. Cormorant 16: 34-40.

Brooke, R.K., Cooper, J., Hockey, P.A.R., Ryan, P.G., Sinclair, J.C., Suter, W. and Tree, A.J. 1988. Distribution, population size and conservation of the Antarctic Tern Sterna vittata in southern Africa. Cormorant 16: 107-113.

Fraser, M.W., Ryan, P.G. and Watkins, B.P. 1988. The seabirds of Inaccessible Island, South Atlantic Ocean. Cormorant 16: 7-33.

Ryan, P.G. 1987. The distribution, population size and foraging behaviour of Kerguelen Terns at the Prince Edward Islands. South African Journal of Antarctic Research 17: 163-166.

Seabird-fishery interactions (47 papers)

Masiko, O.B., Ryan, P.G., van der Lingen, C.D., Upfold, L., Somhlaba, S., Masotla, M., Geja, Y., Dyer, B.M., Crawford, R.J.M. and Makhado, A.B. 2021. Are Cape Cormorants Phalacrocorax capensis losing the competition? Dietary overlap with commercial fisheries. Ostrich 92: 280-294.

Grémillet, D., Collet, J., Weimerskirch, H., Courbin, N., Ryan, P.G. and Pichegru, L. 2019. Bird-borne ship radar detectors show surprisingly few vessel encounters in Cape gannets competing with fisheries. PLoS One 14(2): e0210328.

Rollinson, D.P., Wanless, R.M. and Ryan, P.G. 2017. Patterns and trends in seabird bycatch by the pelagic longline fishery off South Africa. African Journal of Marine Science 39: 9-25.

Sherley, R.B., Botha, P., Underhill, L.G., Ryan, P.G., van Zyl, D., Cockroft, A.C., Crawford, R.J.M., Dyer, B.M. and Cook, T.R. 2017. Defining ecologically-relevant scales for spatial protection using long-term data on an endangered seabird and local prey availability. Conservation Biology 31: 1312-1321.

Burg, T., Catry, P., Ryan, P.G. and Phillips, R.A. 2017. Genetic population structure of Black-browed and Campbell Albatrosses, and implications for assigning provenance of birds killed in fisheries. Aquatic Conservation: Marine and Freshwater Ecosystems 27: 1156-1163.

Schoombie, S., Dilley, B.J., Davies, D., Glass, T. and Ryan, P.G. 2017. The distribution of breeding Sooty Albatrosses from the three most important breeding sites: Gough, Tristan and the Prince Edward Islands. Emu 117: 160-169.

Rollinson, D.P., Dilley, B.J. and Ryan, P.G. 2016. Diving behaviour of Grey Petrels and its relevance for mitigating longline bycatch. Emu 116: 240-249.

Crawford, R.J.M., Randall, R.M., Cook, T.R., Ryan, P.G., Dyer, B.M., Fox, R., Geldenhuys, D., Huisamen, J., McGeorge, C., Smith, M.K., Upfold, L., Visagie, J., Whittington, P.A., Wilke, C.G. and Makhado, A.B. 2016. Cape cormorants decrease, move east and adapt foraging strategies following eastward displacement of their main prey. African Journal of Marine Science 38: 373-383.

Grémillet, D., Péron, C., Kato, A., Amélineau, F., Ropert-Coudert, Y., Ryan, P.G. and Pichegru, L. 2016. Starving seabirds: Unprofitable foraging and its fitness consequences in Cape gannets competing with fisheries in the Benguela upwelling ecosystem. Marine Biology 163: 35.

Techow, N.M.S.M., O'Ryan, C., Robertson, C.J.R. and Ryan, P.G. 2016. The origins of white-chinned petrels killed by long-line fisheries off South Africa and New Zealand. Polar Research 35: 21150.

Rollinson, D.P., Dilley, B.J. and Ryan, P.G. 2014. Diving behaviour of white-chinned petrels and its relevance for mitigating longline bycatch. Polar Biology 37: 1301-1308.

Glass, J.P. and Ryan, P.G. 2013. Reduced seabird night strikes and mortality in the Tristan rock lobster fishery. African Journal of Marine Science 35: 589-592.

Reid, T.A., Wanless, R.M., Hilton, G.M., Phillips, R.A. and Ryan, P.G. 2013. Foraging range and habitat associations of non-breeding Tristan Albatrosses: overlap with fisheries and implications for conservation. Endangered Species Research 22: 39-49.

Tew Kai, E., Benhamou, S., van der Lingen, C., Coetzee, J., Pichegru, L., Ryan, P.G. and Grémillet, D. 2013. Are Cape gannets dependent upon fishery waste in the southern Benguela upwelling system? A multi-scale analysis using seabird GPS-tracking, hydro-acoustic estimates of pelagic fish, and vessel monitoring systems. Journal of Applied Ecology 50: 659-670.

Pichegru, L., Ryan, P.G., van Eeden, R., Reid, T., Grémillet, D. and Wanless, R. 2012. Industrial fishing, no-take zones and endangered penguins. Biological Conservation 156: 117-125.

Ryan, P.G., Pichegru, L. and Grémillet, D. 2010. Parlous conservation status of African Penguins provides the correct wider context. Biology Letters http://rsbl.royalsocietypublishing.org/content/early/2010/02/04/rsbl.2009.0913.e-letters#

Grémillet, D., Mullers, R.H.E., Moseley, C., Pichegru, L., Coetzee, J.C., Sabarros, P.S., van der Lingen, C., Ryan, P.G., Kato, A. and Ropert-Coudert, Y. 2010. Seabirds, fisheries and cameras. Frontiers in Ecology and the Environment 8: 401-402.

Pichegru, L., Grémillet, D., Crawford, R.J.M., and Ryan, P.G. 2010. Marine no-take zone rapidly benefits threatened penguin. Biology Letters 6: 498-501.

Petersen, S.L., Honig, M.B., Nel, R., Ryan, P.G. and Underhill, L.G. 2009. Turtle bycatch in the pelagic longline fishery off southern Africa. African Journal of Marine Science 31: 87-96.

Petersen, S.L., Honig, M.B., Ryan, P.G. and Underhill, L.G. 2009. Seabird bycatch in the pelagic longline fishery off southern Africa. African Journal of Marine Science 31: 191-204.

Petersen, S.L., Honig, M.B., Ryan, P.G., Underhill, L.G. and Goren, M. 2009. Seabird bycatch in the demersal longline fishery off southern Africa. African Journal of Marine Science 31: 205-214.

Petersen, S.L., Honig, M.B., Ryan, P.G., Underhill, L.G. and Compagno, L.J.V. 2009. Pelagic shark bycatch in the tuna- and swordfish-directed longline fishery off southern Africa. African Journal of Marine Science 31: 215-225.

Pichegru, L., Ryan, P.G., Le Bohec, C., van der Lingen, C.D., Navarro, R., Petersen, S., Lewis, S., van der Westhuizen, J. and Grémillet, D. 2009. Overlap between vulnerable top predators and fisheries in the Benguela upwelling system: implications for marine protected areas. Marine Ecology Progress Series 391: 199-208.

Grémillet, D., Pichegru, L., Kuntz, G., Woakes, A.G., Wilkinson, S., Crawford, R.J.M. and Ryan, P.G. 2008. A Junk-food Hypothesis for gannets feeding on fishery waste. Proceedings of the Royal Society London B 275: 1149-1156.

Ryan, P.G. and Watkins, B.P. 2008. Estimating seabird bycatch by fisheries and its impact on seabird populations. Animal Conservation 11: 260-262.

Petersen, S.L., Phillips, R., Ryan, P.G. and Underhill, L.G. 2008. Albatross overlap with fisheries in the Benguela Upwelling System: implications for conservation and management. Endangered Species Research 5: 117-127.

Watkins, B.P., Petersen, S.L. and Ryan, P.G. 2008. Interactions between seabirds and deep-water hake trawl gear: an assessment of impacts in South African waters. Animal Conservation 11: 247-254.

Baker, G.B., Double, M.C., Gales, R., Tuck, G.N., Abbott, C.L., Ryan, P.G., Petersen, S.L, Robertson, C.J.R. and Alderman, R. 2007. A global assessment of the impact of fisheries-related mortality on Shy and White-capped albatrosses: conservation implications. Biological Conservation 137: 319-333.

Abbott, C.L., Double, M.C., Gales, R., Baker, G.B., Lashko, A., Robertson, C.J.R. and Ryan, P.G. 2006. Molecular provenance analysis for Shy and White-capped Albatrosses killed by fisheries interactions in Australia, New Zealand and South Africa. Conservation Genetics 7: 531-542.

Ryan, P.G., Nel, D.C. and Cooper, J. 2006. Reducing longline mortality of seabirds: a South African perspective. Acta Zoologica Sinica 52S: 645-648.

Cuthbert, R.J., Hilton, G., Ryan, P.G. and Tuck, G. 2005. At-sea distribution of the Tristan Albatross Diomedea dabbenena and potential interactions with pelagic longline fishing in the South Atlantic Ocean. Biological Conservation 121: 345-355.

Mills, M.S.L. and Ryan, P.G. 2005. Modelling impacts of long-line fishing: what are the effects of pair-bond disruption and sex-biased mortality on albatross fecundity? Animal Conservation 8: 359-367.

Nel, D.C., Taylor, F., Ryan, P.G. and Cooper, J. 2003. Population dynamics of Wandering Albatrosses Diomedea exulans at sub-Antarctic Marion Island: long-line fishing and environmental influences. African Journal of Marine Science 25: 503-517.

Ryan, P.G., Keith, D.G. and Kroese, M. 2002. Seabird bycatch by tuna longline fisheries off southern Africa. South African Journal of Marine Science 24: 103-110.

Ryan, P.G. and Watkins, B.P. 2002. Reducing incidental mortality of seabirds with an underwater longline setting funnel. Biological Conservation 104: 127-131.

Nel, D.C., Ryan, P.G., Nel, J.L., Klages, N.T.W., Wilson, R.P., Robertson, G. and Tuck, G.N. 2002. Foraging interactions between Wandering Albatrosses Diomedea exulans breeding on Marion Island and long-line fisheries in the southern Indian Ocean. Ibis 144: E141-E154.

Nel, D.C., Ryan, P.G. and Watkins, B.P. 2002. Seabird mortality in the Patagonian toothfish longline fishery around the Prince Edward Islands, 1996-2000. Antarctic Science 14: 151-161.

Nel, D.C., Ryan, P.G., Crawford, R.J.M., Cooper, J. and Huyser, O.A.W. 2002. Population trends of albatrosses and petrels breeding at sub-Antarctic Marion Island. Polar Biology 25: 81-89.

Glass, N., Lavarello, I., Glass, J.P. and Ryan, P.G. 2000. Longline fishing at Tristan da Cunha: impacts on seabirds. Atlantic Seabirds 2: 49-56.

Nel, D.C., Nel, J.L., Ryan, P.G., Klages, N.T.W., Wilson, R.P. and Robertson, G. 2000. The foraging ecology of Grey-headed Mollymawks at Marion Island, southern Indian Ocean, in relation to known longline fishing activity. Biological Conservation 96: 219-231.

Ryan, P.G. 1999. Sexual dimorphism, moult and body condition of seabirds killed by longline vessels around the Prince Edward Islands, 1996-97. Ostrich 70: 187-192.

Ryan, P.G. and Boix-Hinzen, C. 1999. Consistent male-biased seabird mortality in the Patagonian Toothfish long-line fishery. Auk 116: 851-854.

Ryan, P.G. and Boix-Hinzen, C. 1998. Tuna longline fisheries off southern Africa: the need to limit seabird bycatch. South African Journal of Science 94: 179-182.

Barnes, K.N., Ryan, P.G. and Boix-Hinzen, C. 1997. The impact of the hake Merluccius spp. longline fishery off South Africa on procellariiform seabirds. Biological Conservation 82: 227-234.

Moloney, C.L., Cooper, J., Ryan, P.G. and Siegfried, W.R. 1994. Use of a population model to assess the impact of longline fishing on Wandering Albatross Diomedea exulans populations. Biological Conservation 70: 195-204.

Ryan, P.G. 1991. The impact of the commercial lobster fishery on seabirds at the Tristan da Cunha islands, South Atlantic Ocean. Biological Conservation 57: 339-350.

Ryan, P.G. and Cooper, J. 1991. Rockhopper Penguins and other marine life threatened by driftnet fisheries at Tristan da Cunha. Oryx 25: 76-79.

Ryan, P.G. and Moloney, C.L. 1988. The effect of trawling on bird and seal distributions in the southern Benguela region. Marine Ecology Progress Series 45: 1-11.

Foraging ecology of seabirds and other marine predators (84 papers)

Reusch, K., Connan, M., Butler, M.J., Ryan, P.G. and Pichegru, L. subm. Spatio-temporal differences in the diet and trophic ecology of Kelp Gulls Larus dominicanus in South Africa. Ibis

Nourami, E., et al. 2023. Seabird morphology determines operational wind speeds, tolerable maxima and responses to extremes. Current Biology 

Bentley, L.K., Manica, A., Ryan, P.G., Dilley, B.J. and Phillips, R.A. 2022. Divergent foraging habitat preferences between summer-breeding and winter-breeding Procellaria petrels. Ibis (doi:10.1111/ibi.13152)

Powers, K.D., Pratte, I., Ronconi, R.A., Wong, S., Ryan, P.G., Welch, L.J., Silva, T., Hatch, K.A., Westgate, A.J. and Wiley, D. 2022. Age-related interactions with wind during migration support the hypothesis of developmental learning in a migrating long-lived seabird. Frontiers in Marine Science 9: 938033 (doi:10.3389/fmars.2022.938033)

Carpenter-Kling, T., de Blocq, A., Hagen, C., Harding, C., Morris, T., Pichegru, L., Roberts, J., Ryan, P.G., Wanless, R. and McInnes, A. 2022. Important marine areas for endangered African penguins before and after the crucial stage of moulting. Scientific Reports 12: 9489 (doi:10.1038/s41598-022-12969-w).

Bonnet-Lebrun, A.-S., Dias, M.P., Phillips, R.A., Granadeiro, J.P., Brooke, M. de L., Chastel, O., Clay, T.A., Fayet, A., Gilg, O., González-Solis, J., Guilford, T., Hanssen, S.A., Hedd, A., Jaeger, A., Krietsch, J., Lang, J., LeCorre, M., Moe, B., Montevecchi, W.A., Peter, H.-U., Pinet, P, Rayner, M., Reid, T., Ryan, P.G., Schmidt, N.M., Thompson, D., van Bremmelen, R., Wakefield, E., Watanuki, Y., Weimerskirch, H., Yamamoto, T. and Catry, P. 2021. Seabird migration strategies: flight budgets, diel activity patterns and the influence of the moon. Frontiers in Marine Science (doi:10.3389/fmars.2021.683071).

Schoombie, S., Connan, M., Dilley, B.J., Davies, D., Makhado, A.B. and Ryan, P.G. 2022. Non-breeding distribution, activity patterns and moulting areas of Sooty Albatrosses (Phoebetria fusca) inferred from geolocators, satellite trackers and biochemical markers. Polar Biology 45: 31–44 (doi:10.1007/s00300-021-02969-3). 

Dakwa, F.E., Ryan, P.G., Dyer, B.M., Crawford, R.J.M., Pistorius, P.A. and Makhado, A.B. 2021. Long-term variation in the breeding diets of Macaroni and Eastern Rockhopper penguins at Marion Island (1994-2018). African Journal of Marine Science 43 (doi:10.2989/1814232X.2021.1901248)

Gaglio, D., Walton, J. and Ryan, P.G. 2021. Prey delivery to a breeding colony of Caspian Terns Hydroprogne caspia on the west coast of South Africa. Ostrich 92: 140-146. (doi:10.2989/00306525.2020.1856211)

Steinfurth, A., Oppel, S., Dias, M.P., Starnes, T., Pearmain, L., Dilley, B.J., Davies, D., Nydegger, M., Bell, C., le Bouard, F., Bond, A.L., Cuthbert, R.J., Glass, T., Makhado, A.B., Crawford, R.J.M., Ryan, P.G., Wanless, R. and Ratcliffe, N. 2020. Important marine areas for the conservation of northern rockhopper penguins breeding in the Tristan da Cunha archipelago. Endangered Species Research 43: 409–420.

Carpenter-Kling, T., Reisinger, R.R., Orgeret, F., Connan, M., Stevens, K., Ryan, P.G., Makhado, A. and Pistorius, P.A. 2020. Response of four sympatric sub-Antarctic albatross species to environmental variability. Ecology and Evolution

Reusch, K., Suárez, N., Ryan, P.G. and Pichegru, L. 2020. Foraging movements of breeding Kelp Gulls in South Africa. Movement Ecology 8(1). (doi:10.1186/s40462-020-00221-x)

Courbin, N., Chinho, T., Pichegru, L., Verma-Grémillet, A., Péron, C., Ryan, P.G., Grémillet, D. 2020. The dance of the Cape gannet contains social information on foraging behaviour. Animal Behaviour 166: 95–108.

Hindell, M.A., Reissinger, R.R., Ropert-Coudert, Y., et al. 2020. Tracking of marine predators to protect Southern Ocean ecosystems. Nature 580: 87–92. (doi:10.1038/s41586-020-2126-y)

Carneiro A.P.B., Pearmain, E., Oppel, S., Clay, T., Phillips, R.A., Bonnet-Lebrun, A-S., Wanless, R., Abraham, E., Richard, Y., Rice, J., Handley, J., Davies, T., Dilley, B.J., Ryan, P.G., Small, C., Arata, J., Arnould, J., Bell, E., Bugoni, L., Campioni, L., Catry, P., Cleeland, J., Deppe, L., Elliott, G., Freeman, A., Gonzalez-Solis, J., Granadeiro, J.P., Grémillet, D,. Landers, T., Makhado, A., Nel, D., Nicholls, D., Rexer-Huber, K., Robertson, C.J., Sagar, P., Scofield, P., Stahl, J-C., Stanworth, A., Stevens, K., Trathan, P., Thompson, D., Torres, L., Walker, K., Waugh, S., Weimerskirch, H., Dias, M. 2020. A framework for mapping the distribution of seabirds by integrating tracking, demography and phenology. Journal of Applied Ecology  (doi:10.1111/1365-2664.13568)

Requena, S., Oppel, S., Bond, A.L., Hall, J., Cleeland, J., Crawford, R.J.M., Davies, D., Dilley, B.J., Glass, T., Makhado, A.B., Ratcliffe, N., Reid, T.A., Ronconi, R.A., Schofield, A., Steinfurth, A., Wege, M., Beste, M.N. and Ryan, P.G. 2020. Marine hotspots of activity inform protection of a threatened community of pelagic species in a large oceanic jurisdiction. Animal Conservation 23: 585–596. (doi:10.1111/acv.12572)

Jones, C.W., Phillips, R.A., Grecian, W.J. and Ryan, P.G. 2020. Ecological segregation of two superabundant, morphologically-similar, sister seabird taxa breeding in sympatry. Marine Biology 167: 45. (doi:10.1007/s00227-020-3645-7)

Pastor-Prieto, M., Ramos, R., Reyes-González, J.M., Rivas, M.L., Ryan, P.G. and González-Solís, J. 2019. Phenological variability in the spatial ecology of the Endangered Atlantic Petrel, Pterodroma incerta. Endangered Species Research 40: 189-206. (doi:10.3354/esr00991)

Schoombie, S., Schoombie, J., Brink, C.W., Stevens, K.L., Jones, C., Risi, M.M. & Ryan, P.G. 2019. Automated extraction of bank angles from bird-borne video footage using open-source software. Journal of Field Ornithology (doi:10.1111/jofo.12313)

McInnes, A.M., Ryan, P.G., Lacerda, M. and Pichegru, L. 2019. Targeted prey fields determine foraging effort thresholds of a marine diver: important cues for the sustainable management of fisheries. Journal of Applied Ecology (doi:10.1111/1365-2664.13462)

Oppel, S., Bolton, M., Carneiro A.P.B., and 68 others. 2018. Spatial scales of marine conservation management for breeding seabirds. Marine Policy 98: 37-46.

Schoombie, S., Dilley, B.J., Davies, D. and Ryan, P.G. 2018. The foraging range of Great Shearwaters (Ardenna gravis) breeding on Gough Island. Polar Biology 41: 2451-2458. (doi:10.1007/s00300-018-2381-7)

Gaglio, D., Sherley, R.B., Cook, T.R., Ryan, P.G. and Flower, T. 2018. The costs of kleptoparasitism: a non-invasive study of mixed-species seabird breeding colonies. Behavioral Ecology 29: 939-947.

Makhado, N., Crawford, R.J.M., Dias, M.P., Lamont, T., Reisinger, R.R., Dyer, B.M., Pistorius, P., Ryan, P.G., Upfold, L. and Weimerskirch, H. 2018. Foraging behaviour and habitat use by Indian Yellow-nosed Albatrosses Thalassarche carteri breeding at Prince Edward Island. Emu 118: 353-362.

Gaglio, D., Sherley, R.B., Ryan, P.G. and Cook, T.R. 2018. A non-invasive approach to estimate the energetic requirements of an increasing seabird population in a perturbed marine ecosystem. Scientific Reports 8: 8343. (doi:10.1038/s41598-018-26647-3)

Gaglio, D., Cook, T.R., Sherley, R.B. and Ryan, P.G. 2018. How many can you catch? Factors influencing the occurrence of multi-prey loading in provisioning Greater Crested Terns. Ostrich (doi:10.2989/00306525.2018.1439116)

Rollinson, D.P., Dilley, B.J., Davies, D. and Ryan, P.G. 2018. Year-round movements of white-chinned petrels from Marion Island, southwestern Indian Ocean. Antarctic Science 30: 183-195. (doi:10.1017/S0954102018000056)

Ronconi R.R., Schoombie, S., Koopman, H., Westgate, A., Wong, S. and Ryan, P.G. 2018. Effects of age, sex, colony and breeding phase on marine space use by Great Shearwaters Ardenna gravis in the South Atlantic. Marine Biology 165: 58 (doi:10.1007/s00227-018-3299-x)

Gaglio, D., Cook, T.R., McInnes, A., Sherley, R.B., and Ryan, P.G. 2018. Foraging plasticity in seabirds: a non-invasive study of the diet of greater crested terns breeding in the Benguela Region. PLoS One 13(1): e0190444.

Reisinger, R.R., Raymond, B., Hindell, M.A., Bester, M.N., Crawford, R.J.M., Davies, D., de Bruyn, P.J.N., Dilley, B.J., Kirkman, S.P., Makhado, A.B., Ryan, P.G., Schoombie, S., Stevens, K., Sumner, M.D., Tosh, C.A., Wege, M., Whitehead, T.O., Wotherspoon, S. and Pistorius, P.A. 2018. Habitat modelling of tracking data from multiple marine predators identifies important areas in the Southern Indian Ocean. Diversity and Distributions 24: 535-550. (doi:10.1111/ddi.12702)

Ramos, R., Carlile, N., Madeiros, J., Ramírez, I., Paiva, V.H., Dinis, H., Zino, F., Biscoito, M., Leal, G.R., Bugoni, L., Jodice, P.G.R., Ryan, P.G. and González-Solís, J. 2017. It is the time for oceanic seabirds: tracking year-round distribution of gadfly petrels across the Atlantic Ocean. Diversity and Distributions 23: 794-805.

Whitehead, T.O., Connan, M., Ropert-Coudert, Y. and Ryan, P.G. 2017. Subtle but significant segregation in the feeding ecology of sympatric penguins during the critical pre-moult period. Marine Ecology Progress Series 565: 227–236.

Cook, T.R., Gubiani, R., Ryan, P.G. and Muzaffar, S.B. 2017. Group foraging in Socotra cormorants: a bio-logging approach to the study of a complex behavior. Ecology and Evolution 7: 2025-2038.

Gaglio, D., Cook, T.R., Connan, M., Ryan, P.G. and Sherley, R.B. 2017. Dietary studies in birds: testing a non-invasive method using digital photography in seabirds. Methods in Ecology and Evolution 8: 214-222.

McInnes, A.M., Ryan, P.G., Lacerda, M., Deshayes, J., Goschen, W.S., Pichegru, L. 2017. Small pelagic fish responses to fine-scale oceanographic conditions: implications for the endangered African Penguin. Marine Ecology Progress Series 569: 187-203.

Clay, T., Manica, A., Ryan, P.G., Silk, J., Croxall, J.P., Ireland, L., Phillips, R.A. 2016. Proximate drivers of spatial segregation in non-breeding albatrosses. Scientific Reports 6: 29932.

van Eeden, R., Reid, T.A., Ryan, P.G. and Pichegru, L. 2016. Fine-scale foraging cues for African Penguins in a highly variable marine environment. Marine Ecology Progress Series 543: 257–271.

Whitehead, T.O., Kato, A., Ropert-Coudert, Y. and Ryan, P.G. 2016. Habitat use and diving behaviour of Macaroni Eudyptes chrysolophus and Eastern Rockhopper E. chrysocome filholi penguins during the critical pre-moult period. Marine Biology 163: 19.

Grecian, J., McGill, R., Phillips, R.A., Ryan, P.G. and Furness, R.W. 2015. Quantifying variation in δ¹³C and δ¹⁵N isotopes within and between feathers and individuals; is one sample enough? Marine Biology 162: 733-741.

Massie, P., McIntyre, T., Ryan, P.G., Bester, M.N., Bornemann, H. and Ansorge, I. 2015. The role of eddies on the diving behaviour of female Southern Elephant Seals. Polar Biology 39: 297-307.

McInnes, A.M., Khoosal, A., Murrell, B., Merkle, D., Lacerda, M., Nyengera, R., Coetzee, J.C., Edwards, L.C., Ryan, P.G., Rademan, J., van der Westhuizen, J.J. and Pichegru, L. 2015. Recreational fish-finders - an inexpensive alternative to scientific echo-sounders for unravelling the links between marine top predators and their prey. PLoS One 10(11): e0140936.

Torres, L.G., Sutton, P.J., Thomspon, D.R., Delord, K., Weimerskirch, H., Sagar, P.M., Sommer, E., Dilley, B.J., Ryan, P.G. and Phillips, R.A. 2015. Poor transferability of species distribution models for a pelagic predator, the Grey Petrel, indicates contrasting habitat preferences across ocean basins. PLoS One 10(3): e0120014.

Cohen, L.A., Pichegru, L., Grémillet, D., Coetzee, J., Upfold, L. and Ryan, P.G. 2014. A decade of changes in prey availability impact the foraging behaviour and fitness of Cape Gannets. Marine Ecology Progress Series 505: 281-293.

Reid, T.A., Ronconi, R.A., Cuthbert, R.J. and Ryan, P.G. 2014. The summer foraging ranges of adult Spectacled Petrels Procellaria conspicillata. Antarctic Science 26: 23-32.

Barquete, V., Strauss, V. and Ryan, P.G. 2013. Stable isotope turnover in blood and claws: a case study in captive African Penguins. Journal of Experimental Marine Biology and Ecology 448: 121-127.

Dilley, B.J., Davies, D., Connan, M., Cooper, J., de Villiers, M., Swart, L., Vandenabeele, S., Ropert-Coudert, Y. and Ryan, P.G. 2013. Giant petrels as predators of albatross chicks. Polar Biology 36: 761-766.

Ryan, P.G., Edwards, L. and Pichegru, L. 2012. African Penguins Spheniscus demersus, bait balls and the Allee effect. Ardea 100: 89-94.

Cook, T.R., Hamann, M., Pichegru, L., Bonadonna, F., Grémillet, D. and Ryan, P.G. 2012. GPS and time-depth loggers reveal underwater foraging plasticity in a flying diver, the Cape Cormorant. Marine Biology 159: 379-387.

Hamann, M.H., Grémillet, D., Ryan, P.G., Bonadonna, F., van der Lingen, C.D. and Pichegru, L. 2012. A hard-knock life: the foraging ecology of Cape Cormorants amidst shifting prey resources and industrial fishing pressure. African Journal of Marine Science 34: 233-240.

Moseley, C., Grémillet, D., Connan, M., Ryan, P.G., Mullers, R.H.E., van der Lingen, C.D., Coetzee, J.C., Crawford, R.J.M., Sabarros, P., McQuaid, C. and Pichegru, L. 2012. Foraging ecology and ecophysiology of Cape Gannets from colonies with contrasting environments and population dynamics. Journal of Experimental Marine Biology and Ecology 422-423: 29-38.

Pichegru, L., Ropert-Coudert, Y., Kato, A., Takahashi, A., Dyer, B.M. and Ryan, P.G. 2011. Diving patterns of female Macaroni Penguins breeding on Marion Island, South Africa. Polar Biology 34: 945-954.

Wright, K.L.B., Pichegru, L. and Ryan, P.G. 2011. Penguins are attracted to dimethyl sulphide at sea. Journal of Experimental Biology 214: 2509-2511.

Ryan, P.G., Pichegru, L., Ropert-Coudert, Y., Grémillet, D. and Kato, A. 2010. On a wing and a prayer: the foraging ecology of breeding Cape Cormorants. Journal of Zoology London 280: 25-32.

Ronconi, R.A., Ryan, P.G. and Ropert-Coudert, Y. 2010. Diving of Great Shearwaters (Puffinus gravis) in cold and warm water regions of the South Atlantic Ocean. PLoS One 5(11): e15508.

Ryan, P.G. 2009. The seabirds of Conrad Rise, Southern Ocean. Bulletin of the African Bird Club 16: 67-77.

Ropert-Coudert, Y., Daunt, F., Kato, A., Ryan, P.G., Lewis, S., Kobayashi, K., Mori, Y., Grémillet, D. and Wanless, S. 2009. Underwater wingbeats extend depth and duration of plunge dives in gannets. Journal of Avian Biology 40: 380-387.

Ryan, P.G., Sommer, E. and Breytenbach, E. 2008. Giant petrels Macronectes hunting Northern Rockhopper Penguins Eudyptes moseleyi at sea. Ardea 96: 129-134.

Cunningham, G.B., Strauss, V. and Ryan, P.G. 2008. African Penguins (Spheniscus demersus) can detect dimethyl sulphide, a prey-related odour. Journal of Experimental Biology 211: 3123-3127.

Grémillet, D., Lewis, S., Drapeau, L., van der Lingen, C.D., Huggett, J.A., Coetzee, J.C., Verheye, H.M., Daunt, F., Wanless, S. and Ryan, P.G. 2008. Spatial match-mismatch in the Benguela upwelling zone: should we expect chlorophyll and SST to predict marine predator distributions? Journal of Applied Ecology 45: 610-621.

Ryan, P.G. 2007. Diving in shallow water: the foraging ecology of darters (Anhingidae). Journal of Avian Biology 38: 507-514.

Ryan, P.G., Petersen, S.L., Simeone, A. and Grémillet, D. 2007. Diving behaviour of African penguins: do they differ from other Spheniscus penguins? African Journal of Marine Science 29: 153-160.

Pichegru, L., Ryan, P.G., van der Lingen, C.D., Coetzee, J., Ropert-Coudert, Y. and Grémillet, D. 2007. Foraging behaviour and energetics of Cape Gannets Morus capensis feeding on live prey and fishery waste in the Benguela upwelling system. Marine Ecology Progress Series 350: 127-136.

Petersen, S.L., Ryan, P.G. and Grémillet, D. 2006. Is food availability limiting African Penguins at Boulders Beach? A comparison of foraging ecology at mainland and island colonies. Ibis 148: 14-26.

Ropert-Coudert, Y., Wilson, R.P., Grémillet, D., Kato, A., Lewis, S. and Ryan, P.G. 2006. Electrocardiogram recordings in free-ranging gannets reveal minimum difference in heart rate during flapping versus gliding flight. Marine Ecology Progress Series 328: 275-284.

Ryan, P.G., Petersen, S.L., Peters, G. and Grémillet, D. 2004. GPS tracking a marine predator: the effects of precision, resolution and sampling rate on foraging tracks of African Penguins. Marine Biology 145: 215-223.

Ryan, P.G. 2004. Pink-backed Pelicans feeding communally with Great White Pelicans. Ostrich 75: 168.

Grémillet, D., Dell’Omo, G., Ryan, P.G., Peters, G., Ropert-Coudert, Y. and Weeks, S.J. 2004. Offshore diplomacy, or how seabirds mitigate intra-specific competition: A case study based on GPS tracking of Cape gannets from neighbouring breeding sites. Marine Ecology Progress Series 268: 265-279.

Ropert-Coudert, Y., Grémillet, D., Kato, A., Ryan, P.G., Naito, Y. and Le Maho, Y. 2004. A fine-scale time budget of Cape Gannets provides insights into the foraging strategies of coastal seabirds. Animal Behaviour 67: 985-992.

Ropert-Coudert, Y., Grémillet, D., Ryan, P.G., Kato, A., Naito, Y. and Le Maho, Y. 2004. Between air and water: the gannet’s plunge dive. Ibis 146: 281-290.

Keith, D.G., Harck, B.I.B., Ryan, P.G. and Mehlum, F. 2002. Post-breeding dispersal of Northern Giant Petrels Macronectes halli from Marion to Bouvet Islands. Marine Ornithology 30: 31.

Nel, D.C., Lutjeharms, J.R.E., Pakhomov, E.A., Ansorge, I.J., Ryan, P.G. and Klages, N.T.W. 2001. Exploitation of mesoscale oceanographic features by Grey-headed Albatrosses Thalassarche chrysostoma in the southern Indian Ocean. Marine Ecology Progress Series 217: 15-26.

Ryan, P.G. and Nel, D.C. 1999. Foraging behaviour of diving petrels. Emu 99: 72-74.

Wilson, R.P., Hustler, K., Ryan, P.G., Burger, A.E. and Nöldeke, E.C. 1992. Diving birds in cold water: do Archimedes and Boyle determine energetic costs? American Naturalist 140: 179-200.

Ryan, P.G. and Moloney, C.L. 1991. Prey selection and temporal variation in the diet of Subantarctic Skuas at Inaccessible Island, Tristan da Cunha. Ostrich 62: 52-58.

Crawford, R.J.M., Ryan, P.G. and Williams, A.J. 1991. Seabird consumption and production in the Benguela and western Agulhas ecosystems. South African Journal of Marine Science 11: 357-375.

Hockey, P.A.R., Ryan, P.G. and Bosman, A.L. 1989. Age-related intraspecific kleptoparasitism and foraging success of Kelp Gulls Larus dominicanus. Ardea 77: 205-210.

Wilson, R.P., Ryan, P.G. and Wilson, M.-P. 1989. Sharing food in the stomachs of seabirds between adults and chicks - a case for delayed gastric emptying. Comparative Biochemistry and Physiology 94A: 461-466.

Hockey, P.A.R., Bosman, A.L. and Ryan, P.G. 1988. What determines prey selection by Kelp Gulls Larus dominicanus in multispecies mussel communities? Cormorant 16: 103-106.

Ryan, P.G. 1987. The foraging behaviour and breeding seasonality of Hartlaub's Gull Larus hartlaubii. Cormorant 15: 23-32.

Ryan, P.G., Hockey, P.A.R. and Bosman, A.L. 1987. The foraging behavior of Gray Gulls at a sandy beach. Wilson Bulletin 99: 271-273.

Ryan, P.G., Wilson, R.P. and Cooper, J. 1987. Intraspecific mimicry and status signals in juvenile African Penguins. Behavioural Ecology and Sociobiology 20: 69-76.

Wilson, R.P., Ryan, P.G., James, A. and Wilson, M.-P. 1987. Conspicuous coloration may enhance prey capture in some piscivores. Animal Behaviour 35: 1558-1560.

Ryan, P.G. and Jackson, S. 1986. Stomach pumping: is killing seabirds necessary? Auk 103: 427-428.

Jackson, S. and Ryan, P.G. 1986. Differential digestion rates of prey by White-chinned Petrels. Auk 103: 617-619.

Seabird breeding biology (16 papers)

Momberg, M., Ryan, P.G., Hedding, D., Schoombie, J., Goddard, K., Craig, K. and le Roux, P. 2023. Factors determining nest-site selection of surface-nesting seabirds: A case study on the world’s largest pelagic bird, the Wandering Albatross Diomedea exulans. Ibis 165: 190–203. (doi:10.1111/ibi.13111)

Bond, A.L., McClelland G.T.W., Cuthbert, R.J., Glass, T., Repetto, J. and Ryan, P.G. 2020. No change in Atlantic Yellow-nosed Albatross (Thalassarche chlororhynchos) egg size over 160 years. Emu

Dilley, B.J., Davies, D., Schoombie, S., Schoombie, J. and Ryan, P.G. 2019. Burrow wars and sinister behaviour among burrow-nesting petrels at sub-Antarctic Marion Island. Ardea 107: 97-102. (doi.org/10.5253/arde.v107i1.a5)

Keogan, K., Daunt, F., Wanless, S., Phillips, R.A., Walling, C.A., Agnew, P., Ainley, D.G., Anker-Nilssen, T., Ballard, G., Barrett, R.T., Barton, K.J., Bech, C., Becker, P., Berglund, P-A., Bollache, L., Bond, A.L., Bouwhuis, S., Bradley, R.W., Burr, Z.M., Camphuysen, K., Catry, P., Chiaradia, A., Christensen-Dalsgaard, S., Cuthbert, R., Dehnhard, N., Descamps, S., Diamond, T., Divoky, G., Drummond, H., Dugger, K.M., Dunn, M.J., Emmerson, L., Erikstad, K.E., Fort, J., Fraser, W., Genovart, M., Gilg, O., González-Solís, J., Granadeiro, J.P., Grémillet, D., Hansen, J., Hanssen, S.A., Harris, M., Hedd, A., Hinke, J., Igual, J.M., Jahncke, J., Jones, I., Kappes, P.J., Lang, J., Langset, M., Lescroël, A., Lorentsen, S.H., Lyver, P.O., Mallory, M., Moe, B., Montevecchi, W.A., Monticelli, D., Mostello, C., Newell, M., Nicholson, L., Nisbet, I., Olsson, O., Oro, D., Pattison, V., Poisbleau, M., Pyk, T., Quintana, F., Ramos, J.A., Ramos, R., Reiertsen, T.K., Rodríguez, C., Ryan, P., Sanz-Aguilar, A., Schmidt, N.M., Shannon, P., Sittler, B., Southwell, C., Surman, C., Svagelj , W.S., Trivelpiece, W., Warzybok, P., Watanuki, Y., Weimerskirch, H., Wilson, P.R., Wood, A.G., Phillimore, A.B. and Lewis, S. 2018. Global phenological insensitivity to shifting ocean temperatures among seabirds. Nature Climate Change 8: 313-318. (doi.org/10.1038/s41558-018-0115-z)

Jones, M.G.W., Dilley, B.J., Hagens, Q.A., Louw, H., Mertz, E.M., Visser, P. and Ryan, P.G. 2016. Wandering Albatross Diomedea exulans breeding phenology at Marion Island. Polar Biology 40: 1139-1148.

Bond, A.L., McClelland G.T.W., Ryan, P.G. and Glass, T. 2016. Using measurements to predict laying order in harvested Northern Rockhopper Penguin (Eudyptes moseleyi) eggs. Emu 116: 467-471.

Ryan, P.G., Ferreira, C., Perold, V., Osborne, A. and Jones, C.W. 2015. Failure to launch: evidence of protracted parental care in albatrosses. Seabird 28: 48-51.

Witteveen, M., Brown, M. and Ryan, P.G. 2015. Pseudo-egg and exotic egg adoption by Kelp Gulls Larus dominicanus vetula. African Zoology 50: 59-61.

Jones, M.G.W. and Ryan, P.G. 2014. Effects of pre-laying attendance and body condition on long-term reproductive success in Wandering Albatrosses. Emu 114: 137-145.

Jones, M.G.W., Dilley, B.J., Hagens, Q.A., Louw, H., Mertz, E.M., Visser, P. and Ryan, P.G. 2014. The effect of parental age, experience and historic reproductive success on Wandering Albatross (Diomedea exulans) chick growth and survival. Polar Biology 37: 1633-1644.

Ryan, P.G., Dyer, B.M., Martin, A.P., Ward, V.L. and Whittington, P.A. 2013. Supernormal clutches in southern African Kelp Gulls Larus dominicanus vetula. Ostrich 84: 157-160.

Jones, M.G.W., Techow, N.M.S.M. and Ryan, P.G. 2012. Dalliances and doubtful dads: What determines extra-pair paternity in socially monogamous Wandering Albatrosses? Behavioural Ecology and Sociobiology 66: 1213-1224.

Wilson, J.W., Burle, M-H., Cuthbert, R., Stirnemann, R.L. and Ryan, P.G. 2010. Breeding success of Northern Rockhopper Penguins (Eudyptes moseleyi) at Gough Island, South Atlantic Ocean. Emu 110: 137-141.

Ryan, P.G., Cuthbert, R.J. and Cooper, J. 2007. Two-egg clutches among albatrosses. Emu 107: 210-213.

Ryan, P.G., Phillips, R.A., Nel, D.C. and Wood, A.G. 2007. Breeding frequency in Grey-headed Albatrosses. Ibis 149: 45-52.

Ryan, P.G. and Watkins, B.P. 1989. Snow Petrel breeding biology at an inland site in continental Antarctica. Colonial Waterbirds 12: 176-184.

Ryan, P.G. and Hunter, S. 1985. Early breeding of Imperial Cormorants at Prince Edward Island. Cormorant 13: 31-34.

Sundry seabird papers (42 papers)

Schoombie, S., Wilson, R.P. and Ryan, P.G. 2023. Wind driven effects on the fine-scale flight behaviour of dynamic soaring wandering albatrosses. Marine Ecology Progress Series (doi:10.3354/meps14265)

Schoombie, J., Schoombie, S., Connan, M., Jones, C.W., Risi, M., Craig, K.J., Smith, L., Ryan, P.G. and Shephard, E. 2023. Impact of wind on crash-landing mortality in Grey‐headed Albatrosses Thalassarche chrysostoma breeding on Marion Island. Marine Ecology Progress Series (doi:10.3354/meps14292)

Krishnan, K., Garde, B., Bennison, A., Cole, N.C., Cole, E.-L., Darby, J., Elliott, K.H., Fell, A., Gómez-Laich, A., de Grissac, S., Jessopp, M., Lempidakis, E., Mizutani, Y., Prudor, A., Quetting, M., Quintana, F., Robotka, H., Roulin, A., Ryan, P.G., Schalcher, K., Schoombie, S., Tatayah, V., Tremblay, F., Weimerskirch, H., Whelan, S., Wikelski, M., Yoda, K., Hedenström A. and Shepard, E.L.C. 2022. The role of wingbeat frequency and amplitude in flight power. Journal of the Royal Society Interface 19: 20220168 (doi:10.1098/rsif.2022.0168)

Ryan, P.G., Ward, V.L. and Miller, S.M. 2022. First record of a Common Diving Petrel from continental Africa, and a summary of diving petrel distribution in the Southern Ocean. Marine Ornithology 50: 217-220.

Ryan, P.G. and Oppel, S. 2022. Notes on the seabirds of Gough Island and at sea between Gough and Cape Town, March–June 2021. Afrotropical Bird Biology 2: 1-8.

Adekola, E., Makhado, A.B., Dyer, B.M., Upfold, L., Crawford, R.J.M. and Ryan, P.G. 2021. Timing, duration and symmetry of moult in Cape Gannets. Ostrich 92: 295-306 (doi:10.2989/00306525.2021.1988745)

Bonnet-Lebrun, A-S., Dias, M.P., Phillips, R.A., Granadeiro, J.P., Brooke, M. de L., Chastel, O., Clay, T.A., Fayet, A., Gilg, O., Gonzales-Solis, J., Guilford, T., Hanssen, S.A., Hedd, A., Jaeger, A., Krietsch, J., Lang, J., Le Corre, M., Militao, T., Moe, B., Montevecchi, W.A., Peter, H-U., Pinet, P., Rayner, M., Reid, T., Reyes-González, J.M., Ryan, P.G., Sagar, P., Schmidt, N.M., Thompson, D., Van Bemmelen, R., Watanuki, Y., Weimerskirch, H., Yamamoto, T. and Catry, P. 2021. Seabird migration strategies: Flight budgets, diel activity patterns and lunar influence. Frontiers in Marine Science (doi:10.3389/fmars.2021.683071)

Osborne, A.M. and Ryan, P.G. 2021. Using digital photography to study moult extent in breeding seabirds. Ostrich 92: 225-228 (doi:10.2989/00306525.2021.1897699)

Bond, A.L., Taylor, C., Kinchin-Smith, D., Fox, D., Witcutt, E., Ryan, P.G., Loader, S.P. and Weimerskirch, H. 2021. A juvenile Tristan Albatross (Diomedea dabbenena) on land at the Crozet Islands. Polar Biology (doi:10.1007/s00300-020-02786-0)

Ryan, P.G., Dilley, B.J., Risi, M.M., Jones, C.W., Osborne, A., Schofield, A., Repetto, J. and Ratcliffe, N. 2019. Three new seabirds recorded at Tristan da Cunha archipelago. Seabird 32: 122–125.

Cook, T., Martin, R., Roberts, J., Häkkinen, H., Botha, P., Meyer, C., Sparks, E., Underhill, L.G., Ryan, P.G. and Sherley, R. 2020. Parenting in a warming world: thermoregulatory responses to heat stress in an endangered seabird. Conservation Physiology 8 (doi:10.1093/conphys/coz109)

Mondreti, R., Davidar, P., Ryan, P.G., Thiebot, J.B. and Grémillet, D. 2020. Seabird and cetacean occurrence in the Bay of Bengal associated with marine productivity and commercial fishing effort. Marine Ornithology 48: 91–101.

Jones, M.G.W., Techow, N.M.S., Risi, M.M., Jones, C.W., Hagens, Q.A., Taylor, F. and Ryan, P.G. 2020. Hybridisation and cuckoldry between Black-browed and Grey-headed Albatrosses. Antarctic Science 32: 10-14. (doi:10.1017/S0954102019000506)

Gittins, O., Roma, L.G., Valle, R., Abad, F.X., Nofrarías, M., Ryan, P.G., González-Solís, J., Majó, N. 2019. Serological and molecular surveys of influenza A viruses in Antarctic and sub-Antarctic wild birds. Antarctic Science 32: 15-20. (doi:10.1017/S0954102019000464)

Ryan, P.G., Lee, J. and Le Bouard, F. 2020. Moult intensity in Blue Petrels, and the discovery of a key moult site off West Antarctica. Antarctic Science 32: 1-9. (doi:10.1017/S0954102019000427)

Connan, M., Ryan, P.G., Whitehead, T.O., Dilley, B.J., Davies, D. and McQuaid, C. 2019. Multidimensional stable isotope analysis illuminates resource partitioning in a sub-Antarctic island bird community. Ecography 42: 1948-1959.

Roscales, J.L., Vicente, A., Ryan, P.G., González-Solís, J. and Jiménez, B. 2019. Spatial and interspecies heterogeneity in concentrations of perfluoroalkyl substances (PFASs) in seabirds of the Southern Ocean. Environmental Science & Technology 53: 9855-9865.

Risi, M., Jones, C.W., Schoombie, S. and Ryan, P.G. 2019. Plumage and bill abnormalities in albatross chicks on Marion Island. Polar Biology (doi:10.1007/s00300-019-02528-x)

Cerda-Cuéllar, M., Moré, E., Ayats, T., Aguilera, M., Muñoz-González, S., Antilles, N., Ryan, P.G. and González-Solís, J. 2019. Do humans spread zoonotic enteric bacteria in Antarctica? Science of the Total Environment 654: 190-196.

Flood, R.L. and Ryan, P.G. 2018. Age-related timing of primary moult in Shy Albatross Thalassarche cauta (sensu lato) longline casualties from South Africa. Marine Ornithology 46: 201-206.

Vanstreels, R.E., Yabsley, M.J., Swanepoel, L., Stevens, K.L., Carpenter-Kling, T., Ryan, P.G. and Pistorius, P. 2018. Molecular characterization and lesions associated with Diomedenema diomedeae (Aproctoidea: Desmidocercidae) from Grey-headed Albatrosses (Thalassarche chrysostoma) on Subantarctic Marion Island. International Journal for Parasitology: Parasites and Wildlife 7: 155-160.

Payo-Payo, A., Sanz-Aguilar, A., Gaglio, D., Sherley, R., Cook, T., Altwegg, R., and Ryan, P.G. 2018. Survival estimates for the Greater Crested Tern (Thalasseus bergii) in southern Africa. African Journal of Marine Science 40: 43-50.

Moré, E., Ayats, T., Ryan, P.G., Naicker, P.R., Keddy, K.H., Gaglio, D., Witteveen, M. and Cerda-Cuéllar, M. 2017. Seabirds (Laridae) as a source of Campylobacter spp., Salmonella spp. and antimicrobial resistance in South Africa. Environmental Microbiology 19: 4164–4176.

Ryan, P.G., Le Bouard, F. and Lee, J. 2017. Westward range extension of Short-tailed Shearwaters in the Southern Ocean. Polar Biology 40: 2323-2327.

Becker, P., Goutner, V., Ryan, P.G. and González-Solís, J. 2016. Feather mercury concentrations in Southern Ocean seabirds: variation by species, site and time. Environmental Pollution 216: 253-263.

Roscales, J.L., González-Solís, J., Zango, L., Ryan, P.G. and Jiménez, B. 2016. Latitudinal exposure to DDTs, HCB, PCBs, PBDEs and DP in giant petrels (Macronectes spp.) across the Southern Ocean. Environmental Research 148: 285–294.

Jones, C.W., Risi, M.M., Kuntz, W., Ryan, P.G., Steinfurth, A. and Bond, A.L. 2015. Bill deformities in penguins (Spheniscidae): a global review. Marine Ornithology 43: 207–209.

Colabuono, F.I., Barquete, V., Taniguchi, S., Ryan, P.G. and Montone, R.C. 2014. Stable isotopes of carbon and nitrogen in the study of organochlorine contaminants in albatrosses and petrels. Marine Pollution Bulletin 83: 241-247.

Commins, M.L., Ansorge, I. and Ryan, P.G. 2014. Multi-scale factors influencing seabird assemblages in the African sector of the Southern Ocean. Antarctic Science 26: 38-48.

Ryan, P.G. 2013. Moult of flight feathers in darters (Anhingidae). Ardea 101: 177-180.

Ryan, P.G., Rose, B., Carter, M. and Clarke, R.H. 2013. A review of Jouanin’s Petrel records in the eastern Indian Ocean. Ostrich 84: 157-160.

Abad, F.X., Busquets, N., Sánchez, A., Ryan, P.G., Majó, N. and González-Solís, J. 2013. Serological and virological surveys of Influenza A virus in Antarctic and sub-Antarctic penguins. Antarctic Science 25: 339-344.

Ropert-Coudert, Y., Grémillet, D., Gachot-Neveu, H., Lewis, S. and Ryan, P.G. 2005. Seeking dimorphism in monomorphic species: the lure of the gannet’s mask. Ostrich 76: 212-214.

Cuthbert, R.J., Phillips, R.A. and Ryan, P.G. 2003. Separating the Tristan Albatross and the Wandering Albatross using morphometric measurements. Waterbirds 26: 338-344.

Ryan, P.G. 2001. Partial leucism in Whitechinned Petrels. Bird Numbers 10(2): 6-7.

Ryan, P.G., Steele, W.K., Siegfried, W.R. and Vogel, J.C. 1992. Radiocarbon dates of Snow Petrel regurgitations can reveal exposure periods for nunataks in Antarctica. South African Journal of Science 88: 578-580.

Ryan, P.G., Avery, G., Rose, B., Ross, G.J.B., Sinclair, J.C. and Vernon, C.J. 1989. The Southern Ocean seabird irruption to South African waters during winter 1984. Cormorant 17: 41-55.

Ryan, P.G. and Cooper, J. 1989. The distribution and abundance of aerial seabirds in relation to Antarctic krill in the Prydz Bay region, Antarctica, during late summer. Polar Biology 10: 199-209.

Ryan, P.G. and Cooper, J. 1989. Observer precision and bird conspicuousness during counts of birds at sea. South African Journal of Marine Science 8: 271-276.

Hoberg, E.P. and Ryan, P.G. 1989. Ecology of helminth parasitism in Puffinus gravis (Procellariiformes) on the breeding grounds at Gough Island. Canadian Journal of Zoology 67: 220-225.

Duffy, D.C., Ryan, P.G., Wilson, R.P. and Wilson, M.-P. 1988. Spring seabird distribution in the Straits of Magellan. Cormorant 16: 98-102.

Ryan, P.G. and Avery, G. 1987. Wreck of juvenile Blackbrowed Albatrosses on the west coast of South Africa during storm weather. Ostrich 58: 139-140.

Ryan, P.G. 1986. Trematode anklets on procellariiform seabirds from southern Africa and the adjacent Southern Ocean. Cormorant 13: 157-161.

Island biology, restoration and conservation (63 papers)

Dilley, B.J., Davies, D., Ortmann, H., Glass, T. and Ryan, P.G. 2021. Breeding biology of two sympatric Nesospiza finches at Nightingale Island, Tristan da Cunha. Afrotropical Bird Biology 1: 5. 

Dilley, B.J., Davies, D., Ortmann, H., Glass, T. and Ryan, P.G. 2021. Population size and home ranges of Nesospiza wilkinsi finches at Nightingale Island, Tristan da Cunha. Ardea 109: 67-76.

Jones, C.W., Risi, M.M., Osborne, A.M., Ryan, P.G. and Oppel, S. 2021. Mouse eradication is required to prevent local extinction of an endangered seabird on an oceanic island. Animal Conservation 24: 637-645. (doi:10.1111/acv.12670)

Dilley, B.J., Swain, G., Repetto, J. and Ryan, P.G. 2021. Using playback to estimate the distribution and density of the world's smallest flightless bird, the Inaccessible Island Rail. Bird Conservation International 31: 286-292. (doi:10.1017/S0959270920000453)

Dilley, B.J., Davies, D., Glass, T., Bond, A.L. and Ryan, P.G. 2020. Severe impact of introduced scale insects on Island Trees threatens endemic finches at the Tristan da Cunha archipelago. Biological Conservation 251: 108761.

Jones, C.W., Risi, M.M., Osborne, A.M., Parker, G.C., Rexer-Huber, K., le Bouard, F., Cleeland, J.B., Lawrence, K., Kinchin-Smith, D., Witcutt, E., Starnes, T., Bond, A.L., Ryan, P.G. and Oppel, S. 2020. Abundance, distribution and breeding success of the endemic Gough Island Finch Rowettia goughensis between 2009 and 2018. Emu 120: 230-238.

Dilley, B.J., Davies, D., Repetto, J., Swain, G. and Ryan, P.G. 2020. Rats and prions at Tristan da Cunha Island. Ostrich 91: 240-243. (doi:10.2989/00306525.2020.1771622) 

Ryan, P.G., Dilley, B.J., Davies, D., Glass, T. and Abadi, F. 2019. Short-term movement patterns, population estimates and breeding biology of an island endemic bird, the Tristan Thrush. Brazilian Journal of Ornithology 27: 234–241.

Ryan, P.G. and Dilley, B.J. 2019. Intertidal foraging by Tristan Thrushes. Ostrich 90: 179-181.

Holmes, N.D., Spatz, D.R., Oppel, S., Tershy, B., Croll, D.A., Keitt, B., Genovesi, P., Burfield, I.J., Will, D.J., Bond, A., Wegmann, A., Aguirre-Muñoz, A., Raine, A.F., Knapp, C., Hung, C-H., Wingate, D., Hagen, E., Méndez-Sánchez, F., Rocamora, G., Yuan, H-W., Fric, J., Millett, J., Russell, J., Liske-Clark, J., Vidal, E., Jourdan, H., Campbell, K., Springer, K., Swinnerton, K., Gibbons-Decherong, L., Langrand, O., Brooke, M. de L., McMinn, M., Bunbury, N., Oliveira, N., Sposimo, P., Geraldes, P., McClelland, P., Hodum, P., Ryan, P.G., Borroto-Páez, R., Pierce, R., Griffiths, R., Fisher, R.N., Wanless, R., Pasachnik, S., Cranwell, S., Micol, T., and Butchart, S.H.M. 2019. Globally important islands for eradicating invasive mammals to benefit highly threatened vertebrates. PLoS One 14(3): e0212128. (doi.org/10.1371/journal. pone.0212128)

Caravaggi, A., Cuthbert, R.J., Ryan, P.G., Cooper, J. and Bond, A. 2019. The cumulative impacts of introduced house mice on the breeding success of nesting seabirds on Gough Island. Ibis 161: 648-661. (doi.org/10.1111/ibi.12664)

Jones, C.W., Risi, M.M., Cleeland, J. and Ryan, P.G. 2018. First evidence of mouse attacks on adult albatrosses and petrels breeding on sub-Antarctic Marion and Gough Islands. Polar Biology 42: 619-623. (doi.org/10.1007/s00300-018-02444-6)

Brooke, M.de L., Bonnaud, E., Dilley, B.J., Flint, B., Holmes, N.D., Jones, H.P., Provost, P., Rocamora, G., Ryan, P.G., Surman, C. and Buxman, R.T. 2018. Enhancing the value of future island eradications needs improved understanding of past outcomes. Animal Conservation 21: 19-20.

Dilley, B.J., Schoombie, S., Stevens, K., Davies, D., Perold, V., Osborne, A., Schoombie, J., Brink, C.W., Carpenter-Kling, T. and Ryan, P.G. 2018. Mouse predation affects breeding success of burrow-nesting petrels at sub-Antarctic Marion Island. Antarctic Science 30: 93-104.

Schoombie, S., Schoombie, J., Oosthuizen, A., Suleman, E., Jones, M.G.W., Pretorius, L., Dilley, B.J. and Ryan, P.G. 2018. Avian pox in seabirds on Marion Island, southern Indian Ocean. Antarctic Science 30: 3-12.

Brooke, M.de L., Bonnaud, E., Dilley, B.J., Flint, B., Holmes, N.D., Jones, H.P., Provost, P., Rocamora, G., Ryan, P.G., Surman, C. and Buxman, R.T. 2017. Seabird population changes following vertebrate eradications on islands. Animal Conservation 21: 3-12.

Rodríguez, A., Holmes, N.D., Ryan, P.G., Wilson, K.-J., Faulquier, L., Murillo, Y., Raine, A.F., Penniman, J.F., Neves, V., Rodríguez, B., Negro, J.J., Chiaradia, A., Dann, P., Anderson, T., Metzger, B., Shirai, M., Deppe, L., Wheeler, J., Hodum, P., Gouveia, C., Carmo, V., Carreira, G.P., Delgado-Alburqueque, L., Guerra-Correa, C., Couzi, F.-X., Travers, M., Le Corre, M. 2017. A global review of seabird mortality caused by land-based artificial lights. Conservation Biology 31: 986-1001.

Bond, A.L., Risi, M.M., Jones, C.W. and Ryan, P.G. 2016. Potential for primary poisoning of a critically endangered endemic land bird during rodent eradication operations at Gough Island, Tristan da Cunha. PeerJ 4: e1744.

Cerfonteyn, M. and Ryan, P.G. 2016. Have burrowing petrels recovered on Marion Island two decades after cats were eradicated? Evidence from Subantarctic Skua prey remains. Antarctic Science 28: 51-57.

Cuthbert, R.J., Wanless, R.M., Angel, A., Burlé, M.-H., Hilton, G.M., Louw, H., Visser, P., Wilson, J.W. and Ryan, P.G. 2016. Drivers of predatory behaviour and extreme body size in House Mice Mus musculus on Gough Island. Journal of Mammalogy 97: 533–544.

Dilley, B.J., Schramm, M. and Ryan, P.G. 2016. Modest increases in densities of burrow-nesting petrels following the removal of cats (Felis catus) from Marion Island Polar Biology 40: 625-637.

Dilley, B.J., Schoombie, S., Schoombie, J. and Ryan, P.G. 2016. ‘Scalping’ of albatross fledglings by introduced mice spreads rapidly at Marion Island. Antarctic Science 28: 73-80.

Davies, D., Dilley, B.J., Bond, A.L., Cuthbert, R.J. and Ryan, P.G. 2015. Trends and tactics of mouse predation on Tristan Albatross Diomedea dabbenena chicks at Gough Island, South Atlantic Ocean. Avian Conservation and Ecology 10(1): 5.

Dilley, B.J., Davies, D., Bond, A.L. and Ryan, P.G. 2015. Effects of mouse predation on burrowing petrel chicks at Gough Island. Antarctic Science 27: 543-553.

Gray, M.M., Parmenter, M., Hogan, C., Ford, I., Cuthbert, R.J., Ryan, P.G., Broman, K.W. and Payseur, B.A. 2015. Genetics of rapid and extreme size evolution in island mice. Genetics 201: 213-228.

Ryan, P.G., Ortmann, H.E. and Herian, K. 2014. The impact of introduced scale insects on Phylica arborea and the implications for Nesospiza finches at Tristan da Cunha. Biological Conservation 176: 48-53.

Cuthbert, R.J., Broome, K., Bradley, J. and Ryan, P.G. 2014. Evaluating the effectiveness of aerial baiting operations for rodent eradications on cliffs on Gough Island, Tristan da Cunha. Conservation Evidence 11: 25-28.

Gray, M.M., Wegmann, D., Haasl, R.J., White, M.A., Gabriel, S.I., Searle, J.B., Cuthbert, R.J., Ryan, P.G. and Payseur, B.A. 2014. Demographic history of a recent invasion of House Mice on the isolated island of Gough. Molecular Ecology 23: 1923-1939.

Cuthbert, R.J., Louw, H., Lurling, J., Parker, G., Rexer-Huber, K., Sommer, E., Visser, P. and Ryan, P.G. 2013. Low burrow occupancy and breeding success of burrowing petrels at Gough Island: a consequence of mouse predation. Bird Conservation International 23: 113-124.

Cooper, J., Cuthbert, R.J. and Ryan, P.G. 2013. An overlooked biosecurity concern: back-loading at islands supporting introduced rodents. Aliens 33: 28-31.

Le Roux, P.C., Ramaswielaa, T., Kalwijb, J.M., Shaw, J.D., Ryan, P.G., Treasure, A.M., McClelland, G.T.W., McGeoch, M.A. and Chown, S.L. 2013. Human activities, propagule pressure and alien plants in the sub-Antarctic: tests of generalities and evidence in support of management. Biological Conservation 161: 18-27.

Ryan, P.G., Glass, J.P., Glass, T., Barendse, J. and Cuthbert, R.C. 2012. Eradication of New Zealand flax Phormium tenax on Inaccessible and Nightingale Islands, Tristan da Cunha. Conservation Evidence 9: 58-62.

Chown, S.L., Lee, J.E., Hughes, K.A., Barnes, J., Barrett, P.J.,Bergstrom, D.M., Convey, P., Cowan, D.A., Crosbie, K., Dyer, G., Frenot, Y., Grant, S.M., Herr, D., Kennicutt, M.C., Lamers, M., Murray, A., Possingham, H.P., Reid, K., Riddle, M., Ryan, P.G., Sanson, L., Shaw, J.D., Sparrow, M.D., Summerhayes, C., Terauds, A. and Wall, D.H. 2012. Challenges to the future conservation of the Antarctic. Science 337: 158-159.

Parker, G.C., Rexer-Huber, K., Parker, K.A., Ryan, P.G., Techow, N.M.S.M. and Cuthbert, R.J. 2012. Morphometric and genetic sexing in the Gough Moorhen Gallinula comeri. Ringing and Migration 27: 20-25.

Rexer-Huber, K., Parker, G.C., Ryan, P.G. and Cuthbert, R.J. 2012. Ringing injuries in the Gough Moorhen Gallinula comeri. Ringing and Migration 27: 53-56.

Wanless, R.M., Ratcliffe, N., Angel, A., Bowie, B.C., Cita, K., Hilton, G.M., Kritzinger, P., Ryan P.G. and Slabber, M. 2012. Predation by of Atlantic Petrel chicks by house mice on Gough Island. Animal Conservation 15: 472-479.

Ryan, P.G., Glass, N. and Ronconi, R.A. 2011. The plants and birds of Stoltenhoff and Middle Islands, Tristan da Cunha. Polar Record 47: 86-89.

Cuthbert, R.J., Visser, P., Louw, H. and Ryan, P.G. 2011. Palatability and efficacy of rodent baits for eradicating house mice (Mus musculus) from Gough Island, Tristan da Cunha. Wildlife Research 38: 196-203.

Ryan, P.G. and Ronconi, R.A. 2010. The Tristan Thrush Nesocichla eremita as seabird predator. Ardea 98: 247-250.

Jones, M.G.W. and Ryan, P.G. 2010. Evidence of mouse attacks on albatross chicks on sub-Antarctic Marion Island. Antarctic Science 22: 39-42.

Wanless, R.M., Cooper, J., Slabber, M.J. and Ryan, P.G. 2010. Risk assessment of birds foraging terrestrially at Marion and Gough Islands to primary and secondary poisoning by rodenticides. Wildlife Research 37: 524-530.

Wanless, R.M., Ryan, P.G., Altwegg, R., Angel, A., Cooper, J., Cuthbert, R. and Hilton, G.M. 2009. From both sides: dire demographic consequences of carnivorous mice and longlining for the Critically Endangered Tristan albatrosses on Gough Island. Biological Conservation 142: 1710-1718.

Ryan, P.G. 2008. Important Bird Areas: Tristan da Cunha and Gough Island. British Birds 101: 586-606.

Ryan, P.G. 2008. Taxonomic and conservation implications of ecological speciation in Nesospiza buntings at Tristan da Cunha. Bird Conservation International 18: 30–39.

Ryan, P.G. and Cuthbert, R.J. 2008. The biology and conservation status of the Gough Bunting Rowettia goughensis. Bulletin of the British Ornithologists’ Club 128: 242-253.

Wanless, R.M., Parkes, J., Cooper, J., Fisher, P., Ryan, P.G. and Slabber, M. 2008. Bait acceptance by house mice: an island field trial. Wildlife Research 35: 806-811.

Wanless, R.M., Angel, A., Cuthbert, R.J., Hilton, G.M. and Ryan, P.G. 2007. Can predation by invasive mice drive seabird extinctions? Biology Letters 3: 241-244.

de Villiers, M.S., Cooper, J. and Ryan, P.G. 2005. Individual variability of behavioural responses by Wandering Albatrosses (Diomedea exulans) to human disturbance. Polar Biology 28: 255-260.

Ryan, P.G., Smith, V.R. and Gremmen, N.J.M. 2003. The distribution and spread of alien vascular plants on Prince Edward Island. African Journal of Marine Science 25: 555-561.

Jones, A.G., Chown, S.L., Ryan, P.G., Gremmen, N.J.M. and Gaston, K.J. 2003. A review of conservation threats on Gough Island: a case study for terrestrial conservation in the Southern Oceans. Biological Conservation 113: 75-87.

Ryan, P.G. and Moloney, C.L. 2002. Breeding behaviour, clutch size and egg dimensions of Nesospiza buntings at Inaccessible Island, Tristan da Cunha. Ostrich 73: 52-58.

Ryan, P.G., Cooper, J. and Glass, J.P. 2001. Population status, breeding biology and conservation of the Tristan Albatross Diomedea [exulans] dabbenena. Bird Conservation International 11: 35-48.

Cooper, J. and Ryan, P.G. 2001. The Agreement on the Conservation of Albatrosses and Petrels: implications for research and monitoring at the Prince Edward Islands. South African Journal of Science 97: 78-79.

Huyser, O.A.W., Ryan, P.G. and Cooper, J. 2000. Changes in population size, habitat use and breeding biology of Lesser Sheathbills at Marion Island: impacts of cats, mice and climate change? Biological Conservation 92: 299-310.

Fraser, M.W., Ryan, P.G., Dean, W.R.J., Briggs, D.J. and Moloney, C.L. 1994. Biology of the Tristan Thrush Nesocichla eremita. Ostrich 65: 14-25.

Dean, W.R.J., Milton, S.J., Ryan, P.G. and Moloney, C.L. 1994. The role of disturbance in the establishment of indigenous and alien plants at Inaccessible and Nightingale Islands in the South Atlantic Ocean. Vegetatio 113: 13-24.

Ryan, P.G. 1993. The ecological consequences of an exceptional rainfall event at Gough Island. South African Journal of Science 89: 309-311.

Cooper, J. and Ryan, P.G. 1993. Benign research on a South Atlantic jewel: towards a management plan for Gough Island. George Wright Forum 9 (3-4): 101-112.

Milton, S.J., Ryan, P.G., Moloney, C.L., Cooper, J., Dean, W.R.J. and Medeiros, A.C. Jr. 1993. Disturbance and demography of Phylica arborea (Rhamnaceae) on the Tristan-Gough group of islands. Botanical Journal of the Linnaean Society 111: 55-70.

Roux, J.P., Ryan. P.G., Milton, S.J. and Moloney, C.L. 1992. Vegetation and checklist of Inaccessible Island, central South Atlantic Ocean, with notes on Nightingale Island. Bothalia 22: 93-109.

Ryan, P.G. and Moloney, C.L. 1991. Tristan Thrushes kill adult White-bellied Storm-Petrels. Wilson Bulletin 103: 130-132.

Ryan, P.G., Moloney, C.L. and Watkins, B.P. 1989. Concern about the adverse effect of introduced mice on island tree Phylica arborea regeneration. South African Journal of Science 85: 626-627.

Ryan, P.G., Watkins, B.P. and Siegfried, W.R. 1989. Morphometrics, metabolic rate and body temperature of the smallest flightless bird: the Inaccessible Island Rail. Condor 91: 465-467.

Avian systematics and evolution (36 papers)

Kennedy, M., Salis, A.T., Seneviratne, S.S., Rathnayake, D., Nupen L.J., Ryan, P.G., Volponi, S., Lubbe, P., Lawrence, N.J. and Spencer, H.G. 2023. Phylogeny of the microcormorants, with the description of a new genus. Zoological Journal of the Linnean Society XX: 1-8 (doi 10.1093/zoolinnean/zlad041)

Cole, T.L., Zhou, C. Fang, M., Pan, H., Ksepka, D., Fiddaman, S., Emerling, C., Thomas, D., Bi, X., Fang, Q., Ellegaard, M., Feng, S., Smith, A., Heath, T., Tennyson, A.J.D., Borboroglu, P.G., Wood, J., Haaden, P., Grosser, S., Bost, C.-A., Cherel, Y., Mattern, T., Hart, T., Sinding, M.-H., Shepherd, L., Phillips, R.A., Quillfeldt, P., Masello, J., Bouzat, J., Ryan, P.G., Thompson, D., Ellenberg, U., Dann, P., Miller, G., Boersma, P.D., Zhao, R., Gilbert, T., Yang, H., Zhang, D. and Zhang, G. 2022. Genomic insights into the secondary aquatic transition of penguins. Nature Communications 13: 3912 (doi:10.1038/s41467-022-31508-9).

Rawlence, N.J., Salis, A.T., Spencer, H.G., Waters, J.M., Scarsbrook, L., Mitchell, K.J., Phillips, R.A, Calderón, L., Cook, T.R., Bost, C-A., Dutoit, L., King, T.M., Masello, J.F., Nupen, L.J., Quillfeldt, P., Ratcliffe, N., Ryan, P.G., Till, C.E. and Kennedy, M. 2022. Rapid radiation of Southern Ocean shags in response to receding sea ice. Journal of Biogeography 49: 942-953 (doi:10.1111/jbi.14360).

Masello, J.F., Ryan, P.G., Shepherd, L.D., Quillfeldt, P., Cherel, Y., Tennyson, A.J.D., Alderman, R., Calderón, L., Cole, T., Cuthbert, R.J., Dilley, B.J., Massaro, M., Miskelly, C.M., Navarro, J., Phillips, R.A., Weimerskirch, H. and Moodley, Y. 2022. Independent evolution of intermediate bill widths in a seabird clade. Molecular Genetics and Genomics 297: 183–198. (doi:10.1007/s00438-021-01845-3).

Abeyrama, D.K., Dempsey, Z.W., Ryan, P.G. and Burg, T.M. 2021. Cryptic speciation and population differentiation in the yellow-nosed albatross species complex. Conservation Genetics 22: 757–766. (doi:10.1007/s10592-021-01358-x).

Miller, S.M., Barnes, K.N., Bloomer, P. and Ryan, P.G. 2021. Phylogeography and morphometric variation of the spike-heeled lark Chersomanes albofasciata complex. Ostrich 92: 94–104.

Feng, S., Stiller, J, Deng, Y. et al. 2020. Densely sampling genomes across the diversity of birds increases the power of comparative genomics analyses. Nature 587: 252-257. (doi.org/10.1038/s41586-020-2873-9)

Pan, H., Cole, T.L., Bi, X., Fang, M., Zhou, C., Yang, Z., Ksepka, D.T., Hart, T., Bouzat, J.L., Argilla, L.S., Bertelsen, M.F., Boersma, P.D., Bost, C.-A., Cherel, Y., Dann, P., Fiddaman, S.R., Howard, P., Labuschagne, K., Mattern, T., Miller, G., Parker, P., Phillips, R.A., Quillfeldt, P., Ryan, P.G., Taylor, H., Thompson, D.R., Young, M.J., Ellegard, M.R., Gilbert, M.T.P., Sinding, M.S., Pacheco, G., Shepherd, L.D., Tennyson, A.J.D., Grosser, S., Kay, E., Nupen, L.J., Ellenberg, U., Houston, D.M., Heath, T.A., Reeve, A.H., Johnson, K., Masello, J.F., Stracke, T., McKinley, B., Borboroglu, P.G., Zhang, D-X, Zhang, G. 2020. High-coverage genomes to elucidate the evolution of penguins. Gigascience 8: giz117.

Cole, T.L., Dutoit, L., Dussex, N., Hart, T., Alexander, A., Younger, J.L., Clucas, G.V., Frugone, M.J., Cherel, Y., Cuthbert, R., Ellenberg, U., Fiddaman, S.R., Hiscock, J., Houston, D., Jouventin, P., Mattern, T., Miller, G., Miskelly, C., Nolan, P., Polito, M.J., Quillfeldt, P., Ryan, P.G., Smith, A., Tennyson, A.J.D., Thompson, D., Wienecke, B., Vianna, J.A. and Waters, J.M. 2019. Receding ice drove parallel expansions in Southern Ocean penguins. Proceedings of the National Academy of Sciences of the U.S.A. 116: 26690-26696. (doi:10.1073/pnas.190408116)

Rexer-Huber, K., Veale, A.J., Catry, P., Cherel, Y., McEwan, J.C., Parker, G.C., Phillips, R.A., Ryan, P.G., Stanworth, A.J., van Stijn, T., Thompson, D.R., Waters, J. and Robertson, B.C. 2019. Genomics detects population structure within and between ocean basins in a circumpolar seabird: the white-chinned petrel. Molecular Ecology 28: 4552-2572.

Masello, J.F., Quillfeldt, P., Sandoval-Castellanos, E., Alderman, R., Calderón, L., Cherel, Y., Cole, T.L., Cuthbert, R.J., Marin, M., Massaro, M., Navarro, J., Phillips, R.A., Ryan, P.G., Shepherd, L.D., Suazo, C.G., Weimerskirch, H. and Moodley, Y. 2019. Additive traits lead to feeding advantage and reproductive isolation, promoting homoploid hybrid speciation. Molecular Biology and Evolution 36: 1671-1685. (doi:10.1093/molbev/msz090)

Cole, T.L., Ksepka, D.T., Mitchell, K.J., Tennyson, A.J.D, Thomas, D.B., Pan, H., Zhang, G., Rawlence, N.J., Wood, J.R., Bover, P., Bouzat, J.L., Cooper, A., Fiddaman, S., Hart, T., Miller, G., Ryan, P.G., Shepherd, L.D., Wilmshurst, J.M., Waters, J.M. 2019. Mitogenomes uncover extinct penguin taxa and reveal island formation as a key driver of speciation. Molecular Biology and Evolution 36: 784-797. (doi:10.1093/molbev/msz017)

Stervander, M., Ryan, P.G., Melo, M. and Hansson, B. 2018. The origin of the world's smallest flightless bird, the Inaccessible Island Rail Atlantisia rogersi (Aves: Rallidae). Molecular Phylogenetics and Evolution 130: 92-98.

Robertson, B.C., Stephenson, B.M., Ronconi, R.A., Goldstien, S., Shepherd, L., Tennyson, A., Carlile, N. and Ryan, P.G. 2016. Phylogenetic affinities of the Fregetta storm-petrels are not black and white. Molecular Phylogenetics and Evolution 97: 170-176.

Moodley, Y., Masello, J., Gopi, M., Cole, T., Thali, M., Alderman, R., Cuthbert, R., Marin, M., Massaro, M., Navarro, J., Phillips, R., Ryan, P.G., Suazo, C., Cherel, Y., Weimerskirch, H. and Quillfeldt, P. 2015. Evolutionary factors affecting the cross-species utility of newly developed microsatellite markers in seabirds. Molecular Ecology Resources 15: 1046-1058. (doi:10.1111/1755-0998.12372)

Silva, M.C., Matias, R., Wanless, R.M., Ryan, P.G., Stephenson, B.M., Bolton, M., Ferrand, N. and Coelho, M. 2015. Understanding the mechanisms of anti-tropical divergence in the seabird White-faced Storm-petrel (Procellariiformes: Pelagodroma marina) using a multi-locus approach. Molecular Ecology 24: 3122–3137.

Ryan, P.G., Bourgeois, K., Dromzeé, S. and Dilley, B.J. 2014. The occurrence of two bill morphs of prions Pachyptila vittata on Gough Island. Polar Biology 37: 727-735.

Ryan, P.G., Klicka, L.B., Barker, F.K. and Burns, K.J. 2013. The origin of finches on Tristan da Cunha and Gough Island, central South Atlantic Ocean. Molecular Phylogenetics and Evolution 69: 299-305.

Alstrom, P., Barnes, K.N., Olsson, U., Barker, F.K., Bloomer, P., Kahn, A.A., Qureshi, M.A., Guillaumet, A., Crochet, P.-A. and Ryan, P.G. 2013. Multilocus phylogeny of the avian family Alaudidae (larks) reveals complex morphological evolution, non-monophyletic genera and hidden species diversity. Molecular Phylogenetics and Evolution 69: 1043-1056.

de Hierro, M.D.G.L., Meléon, M. and Ryan, P.G. 2013. Is carrying feathers a sexually selected trait in House Sparrows? Ethology 119: 1-13.

Jansen van Rensburg, A., Bloomer, P., Ryan, P.G. and Hansson, B. 2012. Genetic variation at the major histocompatibility complex (MHCII beta) in Nesospiza buntings: Effects of genetic drift and shared ancestral polymorphism. BMC Evolutionary Biology 12: 143.

Ryan, P.G., Nupen, L., Rose, B. and Sulieman, A.S. 2010. Geographic variation in Socotra Sparrows Passer insularis. Bulletin of the British Ornithologists’ Club 130: 75-82.

Techow, N.M.S.M., O’Ryan, C., Phillips, R.A., Gales, R., Marin, M., Patterson-Fraser, D., Quintana, F., Ritz, M.S., Thompson, D.R., Wanless, R.M., Weimerskirch, H. and Ryan, P.G. 2010. Speciation and phylogeography of giant petrels Macronectes. Molecular Phylogenetics and Evolution 54: 472–487.

Techow, N.M.S.M., Ryan, P.G. and O’Ryan, C. 2009. Phylogeography and taxonomy of White-chinned and Spectacled Petrels. Molecular Phylogenetics and Evolution 52: 25-33.

de Hierro, M.D.G.L. and Ryan, P.G. 2008. Nest defence and egg rejection in the House Sparrow (Passer domesticus) as protection against conspecific brood parasitism. Behaviour 145: 949-964.

Ritz, M.S., Millar, C., Miller, G.D., Phillips, R.A., Ryan, P.G., Sternkopf, V., Liebers-Helbig, D., Peter, H-U. 2008. Phylogeography of the southern skua complex - rapid colonisation of the Southern Hemisphere during a glacial period and reticulate evolution. Molecular Phylogenetics and Evolution 49: 292-303.

Ryan, P.G., Bloomer, P., Moloney, C.L., Grant, T. and Delport, W. 2007. Ecological speciation in South Atlantic island finches. Science 315: 1420-1423.

Delport, W., Grant, T.J., Ryan, P.G. and Bloomer, P. 2006. Isolation of novel microsatellites, and optimization of published Vidua loci, for evolutionary research on Nesospiza buntings. Molecular Ecology Notes 6: 1180-1183.

Beresford, P., Barker, F.K., Ryan, P.G. and Crowe, T.M. 2005. African endemics span the tree of songbirds (Passeri): Molecular systematics of several evolutionary ‘enigmas’. Proceedings of the Royal Society, London B 272: 849-858.

Ryan, P.G., Wright, D., Oatley, G., Wakeling, J., Cohen, C., Nowell, T.L., Bowie, R.C.K., Ward, V. and Crowe, T.M. 2004. Systematics of Serinus canaries and the status of Cape and Yellow-crowned Canaries inferred from mtDNA. Ostrich 75: 288-294.

Ryan, P.G. 2001. Morphological heritability among a hybrid bunting complex: Nesospiza at Inaccessible Island. Condor 103: 429-438.

Ryan, P.G. and Bloomer, P. 1999. The Long-billed Lark complex: a species mosaic in south-western Africa. Auk 116: 194-208.

Ryan, P.G. 1998. The taxonomic and conservation status of the Spectacled Petrel Procellaria conspicillata. Bird Conservation International 8: 223-235.

Ryan, P.G., Hood, I., Bloomer, P., Komen, J. and Crowe, T.M. 1998. Barlow’s Lark: a new species in the Karoo Lark complex of southwest Africa. Ibis 140: 605-619.

Ryan, P.G. and Bloomer, P. 1997. Geographic variation in Red Lark Certhilauda burra plumage, morphology, song and mitochondrial DNA haplotypes. Ostrich 68: 31-36.

Ryan, P.G., Moloney, C.L. and Hudon, J. 1994. Colour variation and hybridization among Nesospiza buntings on Inaccessible Island, Tristan da Cunha. Auk 111: 314-327.

Impacts of energy infrastructure and related structures (16 papers)

Prinsloo, N., Postma, M., Ryan, P.G., Coetzee, M. and de Bruyn, P.J.N. in press. Estimating bird flight height using 3-D photogrammetry. Journal of Zoology 314: 174-186.

Shaw, J.M., Reid, T.A., Gibbons, B.K., Pretorius, M., Jenkins, A.R., Visagie, R., Michael, M.D. and Ryan, P.G. 2021. A large-scale experiment demonstrates line marking reduces power line collision mortality for large terrestrial birds, but not bustards, in the Karoo, South Africa. Ornithological Applications 123: 1-10.(doi.org/10.1093/ornithapp/duaa067)

Lee, A.T.K., Macray, M.B., Ryan, P.G. and Alexander, G.J. 2021. Tortoise mortality along fence lines in the Karoo region of South Africa. Journal of Nature Conservation 59: 125945.

Perold, V., Ralston, S. and Ryan, P.G. 2020. On a collision course? The large diversity of birds killed by wind turbines in South Africa. Ostrich 91: 228-239. (doi.org/10.2989/00306525.2020.1770889)

Jeal, C., Perold, V., Seymour, C., Ralston, S. and Ryan, P.G. 2019. Utility-scale solar energy facilities – effects on invertebrates in an arid environment. Journal of Arid Environments 168: 1-8.

Jeal, C., Perold, V., Ralston, S. and Ryan, P.G. 2019. Impacts of a Concentrated Solar Power trough facility on birds and other wildlife in South Africa. Ostrich 90: 129-137.

Visser, E., Perold, V., Ralston, S., Camiña, A. and Ryan, P.G. 2019. Assessing the impacts of a utility-scale photovoltaic solar energy facility on birds in the Northern Cape, South Africa. Renewable Energy 133: 1285-1294. (doi:10.1016/j.renene.2018.08.106)

Shaw, J.M., Reid, T.A., Schutgens, M., Jenkins, A.R. and Ryan, P.G. 2018. High power line collision mortality of threatened bustards at a regional scale in the Karoo, South Africa. Ibis 160: 431-446.

Shaw, J.M., Jenkins, A.R., Allan, D.G. and Ryan, P.G. 2016. Population size and trends of Ludwig’s Bustard Neotis ludwigii and other large terrestrial birds in the Karoo, South Africa. Bird Conservation International 26: 69-86.

Shaw, J.M. and Ryan, P.G. 2015. Stable isotopes reveal regional movement patterns in an endangered bustard. Austral Ecology 40: 198-205.

Shaw, J.M., van der Merwe, R., van der Merwe, E. and Ryan, P.G. 2015. Winter scavenging rates under power lines in the Karoo, South Africa. South African Journal of Wildlife Research 45: 122-126.

Schutgens, M., Shaw, J.M. and Ryan, P.G. 2014. Collision mortality of large birds on power lines: estimating scavenger and search bias in the Karoo, South Africa. Ostrich 85: 39-45.

Shaw, J.M., Dilley, B.J., Davies, D. and Ryan, P.G. 2014. Trapping and capture myopathy in Ludwig’s Bustard. South African Journal of Wildlife Research 44: 16-23.

Jenkins, A.R., Shaw, J.M., Smallie, J.J., Gibbons, B., Visagie, R. and Ryan, P.G. 2011. Power line collisions in Ludwig’s Bustard Neotis ludwigii: measuring rates and modelling demographic effects in the Karoo, South Africa. Bird Conservation International 21: 303-310.

Shaw, J.M., Jenkins, A.R., Smallie, J.J. and Ryan, P.G. 2010. Modelling power line collision risk for the Blue Crane Anthropoides paradiseus in South Africa. Ibis 152: 590-599.

Shaw, J.M., Jenkins, A.R., Ryan, P.G. and Smallie, J.J. 2010. A preliminary survey of avian mortality on power lines in the Overberg, South Africa. Ostrich 81: 109-113.

Shorebird biology and conservation (13 papers)

Paijmans, D.M., Louwenthal, D., Ryan, P.G. and Hockey, P.A.R. 2015. Temporal and spatial differences in three-egg clutch frequency of the African Black Oystercatcher. Ostrich 86: 35-41.

Ryan, P.G. 2013. Medium-term changes in coastal bird communities in the Western Cape, South Africa. Austral Ecology 38: 251-259.

Ryan, P.G. and Visagie, J. 2008. African Black Oystercatchers feeding in a terrestrial habitat. Ostrich 79: 243.

Ryan, P.G. and Sinclair, I. 2006. Mussulo – an important shorebird wintering site in Angola. Wader Study Group Bulletin 109: 120.

Ryan, P.G., Underhill, L.G., Cooper, J. and Waltner, M. 1988. Waders (Charadrii) and other water birds on the coast, adjacent wetlands and offshore islands of the southwestern Cape Province, South Africa. Bontebok 6: 1-19.

Ryan, P.G., Bosman, A.L. and Hockey, P.A.R. 1988. Notes on the foraging behaviour of Flying and Magellanic Flightless Steamer-ducks. Wildfowl 39: 29-33.

Siegfried, W.R., Hockey, P.A.R., Ryan, P.G. and Bosman, A.L. 1988. Sex and plumage-type ratios of the Lesser Magellan Goose in southern Chile. Wildfowl 39: 15-21.

Hockey, P.A.R., Bosman, A.L. and Ryan, P.G. 1987. The maintenance of polymorphism and cryptic mimesis in the limpet Scurria veriabilis by two species of Cinclodes (Aves: Furnariinae) in central Chile. Veliger 30: 5-10.

Ryan, P.G., Cooper, J., Hockey, P.A.R. and Berruti, A. 1986. Waders (Charadrii) and other water birds on the coast and adjacent wetlands of Natal, 1980-81. Lammergeyer 36: 1-33.

Ryan, P.G. and Cooper, J. 1985. Waders (Charadrii) and other coastal birds of the northwestern Cape Province, South Africa. Bontebok 4: 1-8.

Hockey, P.A.R. and Ryan, P.G. 1985. Waders on rocky shores: does resource depletion lead to competition? South African Journal of Science 81: 698.

Ryan, P.G., Cooper, J. and Stutterheim, C.J. 1984. Waders (Charadrii) and other coastal birds of the Skeleton Coast, South West Africa. Madoqua 14: 71-78.

Ryan, P.G., Cooper, J., Stutterheim, C.J. and Loutit, R. 1984. An annotated list of the birds of the Skeleton Coast Park. Madoqua 14: 79-90.

Other conservation papers (15 papers)

Bates, A.E., Primack, R.B., Giggar, B.S., Bird, T.J., Clinton, M.E., et al. 2021. Global COVID-19 lockdown highlights humans as both threats and custodians of the environment. Biological Conservation 263 (doi:10.1016/j.biocon.2021.109175)

van Velden, J.L., Altwegg, R., Shaw, K. and Ryan, P.G. 2016. Movement patterns and survival estimates of Blue Cranes in the Western Cape. Ostrich 88: 33-43.

van Velden, J.L., Smith, T. and Ryan, P.G. 2016. Cranes and crops: investigating farmer tolerances towards crop-damage by threatened Blue Cranes (Anthropoides paradiseus) in the Western Cape, South Africa. Environmental Management 58: 972-983.

Maphisa, D.H., Donald, P.F., Buchanan, G.M. and Ryan, P.G. 2009. Habitat use, distribution and breeding ecology of the globally threatened Rudd’s Lark and Botha’s Lark in eastern South Africa. Ostrich 80: 19-28.

Lauret-Stepler, M., Bourjea, J., Roos, D., Pelletier, D., Ryan, P.G., Ciccione, S. and Grizel, H. 2007. Reproductive seasonality and trend of Chelonia mydas in the SW Indian Ocean: a 20 year study based on track counts. Endangered Species Research 3: 217-227.

Owino, A.O. and Ryan, P.G. 2007. Recent papyrus swamp habitat loss and conservation implications in western Kenya. Wetlands Ecology and Management 15: 1-12.

Muntifering, J.R., Dickman, A.J., Perlow, L.M., Hruska, T., Ryan, P.G., Marker, L.L. and Jeo, R.M. 2006. Managing the matrix for large carnivores: a novel approach and perspective from cheetah (Acinonyx jubatus) habitat suitability modelling. Animal Conservation 9: 103-112.

Owino, A.O. and Ryan, P.G. 2006. Habitat associations of papyrus specialist birds at three papyrus swamps in western Kenya. African Journal of Ecology 44: 438-443.

Ryan, P.G., Sinclair, I., Cohen, C., Mills, M.S.L., Spottiswoode, C.N. and Cassidy, R. 2004. The conservation status and vocalisations of threatened birds from the scarp forest of Cuanza Sul, Angola. Bird Conservation International 14: 247-260.

Ryan, P.G. and Spottiswoode, C.N. 2003. Long-billed Tailorbirds (Orthotomus moreaui) rediscovered at Serra Jeci, northern Mozambique. Ostrich 74: 141-145.

Ndang'ang'a, P.K., du Plessis, M.A., Ryan, P.G. and Bennun, L.A. 2002. Grassland decline in Kinangop Plateau, Kenya: implications for conservation of Sharpe's Longclaw (Macronyx sharpei). Biological Conservation 107: 341-350.

Ryan, P.G., Bento, C., Cohen, C., Graham, J., Parker V. and Spottiswoode, C. 1999. The avifauna and conservation status of the Namuli Massif, northern Mozambique. Bird Conservation International 9: 315-331.

Joubert, D.F. and Ryan, P.G. 1999. Differences in mammal and bird assemblages between commercial and communal range-lands in the Succulent Karoo, South Africa. Journal of Arid Environments 43: 287-299.

Ryan, P.G., Cowling, R.M. and Costanza, R. 1995. Valuing fynbos ecosystems: is the fynbos worth conserving? South African Journal of Science 91: 572-573.

Ryan, P.G. and Hockey, P.A.R. 1995. Is the Hottentot Buttonquail really endangered? Ostrich 66: 92-93.

Sundry papers (38 papers)

Brooks, M., Rose, S., Altwegg, R., Lee, A.T.K., Nel, H., Ottosson, U., Retief, E., Reynolds, C., Ryan, P.G., Shema, S., Tende, T., Underhill, L.G. and Thomson, R.L. 2022. The African Bird Atlas Project: a description of the project and BirdMap data collection protocol. Ostrich 93: 223-232.

Zeyl, J.N., Snelling, E.P., Connan, M., Basille, M., Clay. T.A., Joo, R., Phillips, R.A., Pistorius, P.A., Ryan, P.G., Snyman, A. and Clusella-Trullas, S. 2022. Aquatic birds have middle ears adapted to amphibious lifestyles. Scientific Reports 12: 5251.

Adekola, O.E., Allan, D.G., Bernitz, Z., Dlungwana, W. and Ryan, P.G. 2021. Extent and symmetry of tail moult in Amur Falcons. Journal of Ornithology 162: 655–667 (doi:10.1007/s10336-021-01874-0)

Rose, S., Suri, J., Brooks, M. and Ryan, P.G. 2020. COVID-19 and citizen science: lessons learned from southern Africa. Ostrich  91: 188-191. (doi:10.2989/00306525.2020.1783589)

Bester, M.N. and Ryan, P.G. 2019. Stalked barnacles on an adult male sub‐Antarctic fur seal at Inaccessible Island, Tristan da Cunha. Polar Biology (doi:10.1007/s00300-019-02560-x)

Bester, M.N. and Ryan, P.G. 2019. Early pupping of a sub-Antarctic fur seal at Inaccessible Island, Tristan da Cunha. Polar Biology (doi:10.1007/s00300-019-02510-7)

Carstens, K.F., Kassanjee, R., Little, R.M., Ryan, P.G. and Hockey, P.A.R. 2019. The effects of weather, group size and type of nest used on the timing of egg-laying in the Southern Ground Hornbill Bucorvus leadbeateri. Journal of Ornithology 160: 633-640 (doi:10.1007/s10336-019-01654-x)

Carstens, K.F., Kassanjee, R., Little, R.M., Ryan, P.G., and Hockey, P.A.R. 2019. Natal dispersal in the Southern Ground Hornbill Bucorvus leadbeateri. Ostrich 90: 119-127 (doi:10.2989/00306525.2019.1590474)

Carstens, K.F., Kassanjee, R., Little, R.M., Ryan, P.G. and Hockey, P.A.R. 2019. Breeding success and population growth of Southern Ground Hornbills Bucorvus leadbeateri in an area supplemented with nest-boxes. Bird Conservation International 29: 627-643 (doi:10.1017/S0959270919000108)

Fraser, C.I., Morrison, A.K., van Sebille, E., Macaya, E.C., Hogg, A.M., Ryan, P.G., Padovan, A., Jack, C., Valdivia, N., Waters, J.M. 2018. Antarctica’s ecological isolation will be broken by storm-driven dispersal and warming. Nature Climate Change 8: 704-708.

Ansorge, I.J., Skelton, P., Bekker, A., de Bruyn, P.J.N., Butterworth, D., Cilliers, P., Cooper, J., Cowan, D.A., Dorrington, R., Fawcett, S., Fietz, S., Findlay, K.P., Froneman, P.W., Grantham, G.H., Greve, M., Hedding, D., Hofmeyr, G.J.G., Kosch, M., le Roux, P., Lucas, M., MacHutchon, K., Meikeljohn, I, Nel, W., Pistorius, P., Ryan, P., Stander, J., Swart, S., Treasure, A., Vichi, M. and van Vuuren, B.J. 2017. Exploring South Africa’s southern frontier: A 20-year vision for polar research through the South African National Antarctic Programme. South African Journal of Science 113: 11-17.

Dagleish, M.P., Ryan, P.G., Girling, S., Ghazali, M. and Bond, A.L. 2017. Clinical pathology of the vulnerable Gough Moorhen (Gallinula comeri). Journal of Comparative Pathology 157: 246-255.

Dagleish, M.P., Ryan, P.G., Girling, S. and Bond, A.L. 2017. Clinical pathology of the critically endangered Gough Bunting (Rowettia goughensis). Journal of Comparative Pathology 156: 264-274.

Fraser, C., Kay, G., du Plessis, M. and Ryan, P.G. 2017. Breaking down the barrier: dispersal across the Antarctic Polar Front. Ecography 40: 235-237.

Williams, H.J., Holton, M.D., Shepard, E.L.C., Largey, N., Norman, B., Ryan, P.G., Duriez, O., Scantlebury, M., Quintana, F., Magowan, E.A., Marks, N.J., Alagaili, A.N., Bennet, N.C. and Wilson, R.P. 2017. Identification of animal movement patterns using tri-axial magnetometry. Movement Ecology 5: 6 (doi:10.1186/s40462-017-0097-x).

Zoghby, B., Little, R.M., Ryan, P.G. and Hockey, P.A.R. 2016. Patterns of roost site selection and use by Southern Ground-Hornbills in the South African lowveld. Ostrich 87: 125-130.

Zoghby, B., Ryan, P.G., Little, R.M., Reid, T. and Hockey, P.A.R. 2015. Seasonal changes in movement and habitat use by Southern Ground-Hornbills in the South African lowveld. Ostrich 89: 87-95.

Bester, M.N., Ryan, P.G., Bester, W.A. and Glass, T. 2014. Vagrant Antarctic Fur Seals at the Tristan da Cunha Islands. Polar Biology 37: 1701-1703.

Ryan, P.G. 2013. The effect of wind direction on flying fish counts. African Journal of Marine Science 35: 585-587.

Treasure, A.M., Moloney, C.L., Bester, M.N., McQuaid, C.D., Findlay, K.P., Best, P.B., Cowan, D.A., de Bruyn, P.J.N., Dorrington, R.A., Fagereng, A., Froneman, P.W., Grantham, G.H., Hunt, B.P.V., Meiklejohn, K.I., Pakhomov, E.A., Roychoudhury, A.N., Ryan, P.G., Smith, V.R., Chown, S.L. and Ansorge, I.J. 2013. South African research in the Southern Ocean: new opportunities and challenges. South African Journal of Science 109. (doi 10.1590/sajs.2013/a009)

Ryan, P.G. and Branch, G.M. 2012. The November 2011 irruption of buoy barnacles Dosima fascicularis in the Western Cape, South Africa. African Journal of Marine Science 35: 157-162.

Crous, P.W., Groenewald, J.Z., Ryan, P.G. and Wingfield, M.J. 2012. Fungal Planet description sheet 147: Seiridium phylicae. Persoonia 29: 186-187.

Cooper, J., Bester, M.N., Chown, S.L., Crawford, R.J.M., Daly, R., Heyns, E., Lamont, T., Ryan, P.G. and Shaw, J.D. 2009. Biological survey of the Prince Edward Islands, December 2008. South African Journal of Science 105: 317-320.

Ryan, P.G., Porter, R.F., Rose, B. and Sulieman, A.S. 2009. Juvenile plumage of the Socotra Bunting Emberiza socotrana. Sandgrouse 31: 109-111.

Best, P.B., Glass, J.P., Ryan, P.G. and Dalebout, M. 2009. Cetacean records from Tristan da Cunha, South Atlantic. Journal of the Marine Biology Association UK 89: 1023-1032.

Bester, M.N., Visagie, J. and Ryan, P.G. 2009. Summer survey of fur seals at Prince Edward Island, Southern Ocean. African Journal of Marine Science 31: 451-455.

Ansorge, I.J., Roman, R., Durgadoo, J.V., Ryan, P.G., Dlamini, L., Gebhardt, Z., Rainier, S., Smith, M., Mtonsti, T. and Lutjeharms, J.R.E. 2008. The first oceanographic survey of the Conrad Rise. South African Journal of Science 104: 333-336.

Ryan, P.G. and Marshall, H.K. 2005. Variation in the wing-clapping display of Clapper Larks. Ostrich 76: 73-77.

Bragg, C.J., Donaldson, J.D. and Ryan, P.G. 2005. Cape porcupines as ecosystem engineers: aspects of density and disturbance in a geophyte-rich semi-arid environment. Journal of Arid Environments 61: 261-275.

Ryan, P.G. 2004. Nest sites and clutch sizes of House Sparrows at Dassen Island, South Africa. Ostrich 75: 165-167.

Bester, M.N., Ryan, P.G. and Dyer, B.M. 2003. Population numbers of fur seals at Prince Edward Island, Southern Ocean African Journal of Marine Science 25: 549-554.

Crawford, R.J.M., Underhill, L.G., Bester, M.N., Cooper, J., Greyling, M.D., Meÿer, M.A., Petersen, S.L., Ryan, P.G. and Wilkinson, W. 2003. The Prince Edward Islands Millennium Expedition. South African Journal of Science 99: 407-409.

Branch, W.R. and Ryan, P.G. 2001. Additions to the Mozambique herpetofauna: two new lizards from the Namuli Massif, Mozambique. Herpetological Review 32: 281-282.

Cooper, J., Siegfried, W.R., Ryan, P.G., Crafford, J.E. and Stock, W.D. 1991. Effects of ornithogenic products on ecosystem structure and functioning: a new South African Biological Antarctic Research Programme. South African Journal of Science 87: 223-226.

Cockroft, V.H., Peddemors, V.M., Ryan, P.G. and Lutjeharms, J.R.E. 1990. Cetaceans associated with Agulhas Current rings in the Southern Ocean. South African Journal of Antarctic Research 20: 64-67.

Dastych, H., Ryan, P.G. and Watkins, B.P. 1990. Notes on Tardigrada from western Dronning Maud Land (Antarctica), with a description of two new species. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 10: 57-66.

Ryan, P.G. and Watkins, B.P. 1989. The influence of physical factors and ornithogenic products on plant and arthropod abundance at an inland nunatak group in Antarctica. Polar Biology 10: 151-160.

Ryan, P.G., Watkins, B.P., Lewis Smith, R.I., Dastych, H., Eicker, A., Foissner, W., Heatwole, H., Miller, W.R. and Thompson, G. 1989. Biological survey of Robertskollen, western Dronning Maud Land: area description and preliminary species list. South African Journal of Antarctic Research 19: 10-20.

Books (14):

Ryan, P.G. 2023. Seabirds of Southern Africa (Second edition). Struik Nature, Cape Town. 200 pp. (ISBN 978 1 77584 847 9). 

Sinclair, I., Hockey, P., Tarboton, W., Rollinson, D., Perrins, N. and Ryan, P. 2020. SASOL Birds of Southern Africa (Fifth edition). Struik Nature, Cape Town. 464 pp. (ISBN 978 1 77007 925 0). Also in Afrikaans (ISBN 978 1 77007 926 7)

Ryan, P.G. 2017. Seabirds of southern Africa. Struik Nature, Cape Town. 160 pp. (ISBN 978 1 77584 519 5).

Tarboton, W. and Ryan, P.G. 2016. Birds of Kruger Park. Struik Nature, Cape Town. 224 pp. (ISBN 978 1 77584 449 5).

Sinclair, I., Hockey, P., Tarboton, W. and Ryan, P. 2011. SASOL Birds of Southern Africa (Fourth edition). Struik Nature, Cape Town. 464 pp. (ISBN 978 1 77007 925 0). Also in Afrikaans (ISBN 978 1 77007 926 7)

Sinclair, I. and Ryan, P.G. 2010. Birds of Africa South of the Sahara. (Second edition). Struik Nature, Cape Town. 768 pp. (ISBN 978 1 77007 623 5)

Terauds, A., Cooper, J., Chown, S.L. and Ryan, P.G. 2010. Marion and Prince Edward: Africa’s southern islands. SunMedia, Stellenbosch. 176 pp. (ISBN 978 1 920338 42 8).

Sinclair, I. and Ryan, P.G. 2009. Complete Photographic Field Guide: Birds of Southern Africa. Struik Nature, Cape Town. 432 pp. (ISBN 978 1 77007 388 3). Also in Afrikaans (ISBN 978 1 77007 589 4)

Ryan, P.G. 2008. The birds and other wildlife of Kirstenbosch. Struik, Cape Town. 56 pp. (ISBN 978 1 77007 725 6)

Ryan, P.G. (ed.) 2007. Field guide to the animals and plants of Tristan da Cunha and Gough Island. Pisces Publications, Newbury. 162 pp. (ISBN 1 874357 33 1)

Hockey, P.A.R., Dean, W.R.J. and Ryan, P.G. (eds). 2005. Roberts’ Birds of Southern Africa (Seventh edition). The Trustees of the John Voelcker Bird Book Fund, Cape Town. 1296 pp. (ISBN 0 620 34053 3)

Sinclair, I. and Ryan, P.G. 2003. Birds of Africa south of the Sahara. Struik, Cape Town. 760 pp. (ISBN 1 86872 857 9)

Ryan, P.G. 2001. Practical birding. Struik, Cape Town. 96 pp. (ISBN 1 86872 608 8)

Hockey, P.A.R., Underhill, L.G., Neatherway, M. and Ryan, P.G. 1989. Atlas of the birds of the southwestern Cape. Cape Bird Club, Cape Town, 236 pp. (ISBN 0 620 14461 0)

Book chapters, management plans and conference proceedings/abstracts (61):

Honorato-Zimmer, D., Weideman, E.A., Pichegru, L., Ryan, P.G. and Thiel, M. 2022. Marine litter and microplastics in the Western Indian Ocean: current knowledge and recommendations. MSA and Nairobi Convention 2022. WIO Science to Policy Platform Series Vol 1 Issue 1. WIOMSA and Nairobi Convention. Pp. 71-82.

Honorato-Zimmer, D., Weideman, E.A., Pichegru, L., Ryan, P.G. and Thiel, M. 2022. A review of the current status of marine litter and microplastics knowledge in the Western Indian Ocean region. United Nations Environment Programme/Nairobi Convention Secretariat and the Western Indian Ocean Marine Science Association (WIOMSA).

Ryan, P.G. 2020. Tristan da Cunha and Gough Island. In: Riddington, R. (ed.) Birds of the UK Overseas Territories. T & AD Poyser, London. Pp. 277-295. (reprinted from British Birds)

Barnado, T., Pichegru, L. and Ryan, P.G. 2020. Macro-litter monitoring along shorelines. In: Barnado, T. and Ribbink, A.J. (eds) African Marine Litter Monitoring Manual. African Marine Waste Network, Sustainable Seas Trust, Port Elizabeth. Pp.17-31.

Barnado, T., Pichegru, L. and Ryan, P.G. 2020. Meso-litter monitoring along shorelines. In: Barnado, T. and Ribbink, A.J. (eds) African Marine Litter Monitoring Manual. African Marine Waste Network, Sustainable Seas Trust, Port Elizabeth. Pp.32-41.

Barnado, T., Ryan, P.G. and Pichegru, L. 2020. Macro-litter monitoring on land. In: Barnado, T. and Ribbink, A.J. (eds) African Marine Litter Monitoring Manual. African Marine Waste Network, Sustainable Seas Trust, Port Elizabeth. Pp.84-96.

GESAMP. 2019. Guidelines for the monitoring and assessment of plastic litter and microplastics in the ocean. (Kershaw, P.J., Turra, A. and Galgani, F. editors), IMO/FAO/UNESCO-IOC/UNIDO/WMO/IAEA/UN/UNEP/UNDP/ISA Joint Group of Experts on the Scientific Aspects of Marine Environmental Protection. Rep. Stud. GESAMP No. 99, 130 pp. (contributors: Eriksen, M., Galgani, F., Hardesty, D., Hassellov, M., Hong, S.H., Joon Shim, W., Kershaw, P., Lusher, A., Mason, S., Ryan, P., Tahir, A., Takada, H., Turra, A., Wilcox, C. and Zhang, W.)

Preston, G.R., Dilley, B.J., Cooper, J., Beaumont, J., Chauke, L.F., Chown, S.L., Devanunthan, N., Dopolo, M., Fikizolo, L., Heine, J., Henderson, S., Jacobs, C.A., Johnson, F., Kelly, J., Makhado, A.B., Marais, C., Maroga, J., Mayekiso, M., McClelland, G., Mphepya, J., Muir, D., Ngcaba, N., Ngcobo, N., Parkes, J.P., Paulsen, F., Schoombie, S., Springer, K., Stringer, C., Valentine, H., Wanless, R.M. and Ryan, P.G. 2019. South Africa works towards eradicating introduced house mice from sub-Antarctic Marion Island: the largest island yet attempted for mice. In: Veitch, C.R., Clout, M.N., Martin, A.R., Russell, J.C. and West, C.J. (eds). Island invasives: scaling up to meet the challenge. Occasional Paper SSC No. 62, 40-46. IUCN, Gland. https://portals.iucn.org/library/sites/library/files/documents/SSC-OP-062-En.pdf

Ryan, P.G. 2018. Plastics and AEWA waterbirds: incidence and impacts. Unpubl. report to the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA). 35 pp.

Ryan, P.G. 2016. Ingestion of plastics by marine organisms. In: Takada, H. and Karapanagioti, H.K. (eds) Hazardous Chemicals Associated with Plastics in the Environment. Springer, Berlin (32 pp). (doi.org/10.1007/698_2016_21)

Ryan, P.G. 2015. A brief history of marine litter research. In: Bergmann, M., Gutow, L. and Klages, M. (eds) Marine Anthropogenic Litter. Springer International Publishing, Cham. Pp. 1-25. (ISBN 978-3-319-16509-7)

Ryan, P. 2015. Birds. In: Griffiths, C., Day, J. and Picker, M. (eds) Freshwater Life. Struik Nature, Cape Town. Pp. 46-63. (ISBN 978 1 77584)

GESAMP (2015). “Sources, fate and effects of microplastics in the marine environment: a global assessment” (Kershaw, P. J., ed.). (IMO/FAO/UNESCO-IOC/UNIDO/WMO/ IAEA/UN/UNEP/UNDP Joint Group of Experts on the Scientific Aspects of Marine Environmental Protection). Rep. Stud. GESAMP No. 90, 96 pp. (contributors: Anderson, A., Andrady, A., Arthur, C., Baker, J., Bouwman, H., Gall, S., Hidalgo-Ruz, V., Köhler, A., Law, K.L., Leslie, H., Kershaw, P., Pahl, S., Potemra, J., Ryan, P., Joon Shim, W., Thompson, R., Takada, H., Turra, A., Vethaak, D. and Wyles, K.)

Shaw, J. and Ryan, P. 2012. Keeping an eye on bird routes. In: Clavaud, R. (ed). Sustainable development in Africa and satellites. Suds-Concepts, France. Pp. 76-77.

Cooper, J., Cuthbert, R.J., Gremmen, N.J.M., Ryan, P.G. and Shaw, J.D. 2011. Earth, fire and water: applying novel techniques to eradicate the invasive plant, procumbent pearlwort Sagina procumbens, on Gough Island, a World Heritage Site in the South Atlantic. In: Veitch, C.R., Clout, M.N. and Towns, D.R. (eds). Island invasives: Eradication and management. IUCN, (International Union for Conservation of Nature), Gland, Switzerland. Pp. 162-165 (ISBN 978 2 8317 1291 8).

Cuthbert, R., Visser, P., Louw, H., Rexer-Huber, K., Parker, G. and Ryan, P. 2011. Preparations for the eradication of mice from Gough Island: results of bait acceptance trials above ground and around cave systems. In: Veitch, C.R., Clout, M.N. and Towns, D.R. (eds). Island invasives: Eradication and management. IUCN, (International Union for Conservation of Nature), Gland, Switzerland. Pp. 47-50 (ISBN 978 2 8317 1291 8).

Jaramillo, A., Rising, J.D., Copete, J.L., Ryan, P.G. and Madge, S.C. 2011. Species accounts of Emberizidae. In: del Hoyo, J., Elliott, A. and Christie, D. (eds) Handbook of the Birds of the World, Vol. 16. Lynx, Barcelona. Pp. 502-683. (ISBN 978 84 96553 78 1).

Ryan, P.G. 2009. Tristan da Cunha and Gough Island. In R. Gillespie and D. Clague (eds) Encyclopedia of Islands University of California Press, Berkeley. Pp. 929-932. (ISBN 978 0 520 25649 1).

Ryan, P.G. and Bester, M.N. 2008. Pelagic predators. In: Chown, S.N. and Froneman, P.W. (eds) The Prince Edwards Archipelago: Land-sea interactions in a changing ecosystem. Sun Media, Stellenbosch. Pp. 121-164. (ISBN 978-1-920109-85-1).

Petersen, S.L., Nel, D.C., Ryan, P.G. and Underhill, L.G. 2008. Understanding and Mitigating Vulnerable Bycatch in southern African Trawl and Longline Fisheries. WWF South Africa Report Series – 2008/Marine/002. 225 pp.

Petersen, S.L., Ryan, P.G. and Underhill, L.G. 2008. Implications of night setting for seabirds and target catches. In: Petersen, S.L., Nel, D.C., Ryan, P.G. and Underhill, L.G. (eds) Understanding and Mitigating Vulnerable Bycatch in southern African Trawl and Longline Fisheries. WWF South Africa Report Series – 2008/Marine/002. Pp. 160-165.

Petersen, S.L., Honig, M.B., Ryan, P.G., Underhill, L.G. and Goren, M. 2008. Gear configurations, line sink rates and seabird bycatch in pelagic longline fisheries. In: Petersen, S.L., Nel, D.C., Ryan, P.G. and Underhill, L.G. (eds) Understanding and Mitigating Vulnerable Bycatch in southern African Trawl and Longline Fisheries. WWF South Africa Report Series – 2008/Marine/002. Pp. 166-177.

Petersen, S.L., Ryan, P.G., Underhill, L.G. and Goren, M. 2008. The use of circle hooks to reduce turtle bycatch and their effect on other vulnerable species. In: Petersen, S.L., Nel, D.C., Ryan, P.G. and Underhill, L.G. (eds) Understanding and Mitigating Vulnerable Bycatch in southern African Trawl and Longline Fisheries. WWF South Africa Report Series – 2008/Marine/002. Pp. 178-184.

Petersen, S.L., Japp, D., Ryan, P.G. and Underhill, L.G. 2008. Is a 5% fin to trunk ratio universally appropriate as a disincentive to catch sharks? In: Petersen, S.L., Nel, D.C., Ryan, P.G. and Underhill, L.G. (eds) Understanding and Mitigating Vulnerable Bycatch in southern African Trawl and Longline Fisheries. WWF South Africa Report Series – 2008/Marine/002. Pp. 185-190.

Petersen, S.L., Honig, M.B., Ryan, P.G. and Underhill, L.G. 2008. Line sink rates and implications for seabirds, target and non-target fish species in the South African demersal longline fishery. In: Petersen, S.L., Nel, D.C., Ryan, P.G. and Underhill, L.G. (eds) Understanding and Mitigating Vulnerable Bycatch in southern African Trawl and Longline Fisheries. WWF South Africa Report Series – 2008/Marine/002. Pp. 191-205.

Ryan, P.G. 2006. Inaccessible Island Seabird Monitoring Manual. RSPB Research Report 16: 1-32. (ISBN 1 901930 70X).

Ryan, P.G. 2006. Family Cisticolidae. In: del Hoyo, J., Elliott, A. and Christie, D. (eds) Handbook of the Birds of the World, Vol. 11. Lynx, Barcelona. Pp. 378-419. (ISBN 978 96553 06 4).

Ryan, P.G., Dean, W.R.J., Madge, S. and Pearson, D. 2006. Species accounts of Cisticolidae. In: del Hoyo, J., Elliott, A. and Christie, D. (eds) Handbook of the Birds of the World, Vol. 11. Lynx, Barcelona. Pp. 420-490. (ISBN 978 96553 06 4).

Ryan, P.G., de Juana, E., Suárez, F., Alström, P. and Donald, P.F. 2004. Species accounts of Alaudidae. In: del Hoyo, J., Elliott, A. and Christie, D. (eds) Handbook of the Birds of the World, Vol. 9. Lynx, Barcelona. Pp. 542-601. (ISBN 978 87334 69 5).

de Juana, E., Suárez, F. and Ryan, P.G. 2004. Family Alaudidae (Larks). In: del Hoyo, J., Elliott, A. and Christie, D. (eds) Handbook of the Birds of the World, Vol. 9. Lynx, Barcelona. Pp. 496-541. (ISBN 978 87334 69 5).

Tuck, G.N., Polacheck, T., Croxall, J.P., Weimerskirch, H., Ryan, P.G., Nel, D.C., Wayte, S. and Bulman, C.M. 2004. Modelling the impact of fishery incidental mortality on three populations of Wandering Albatross. In: Tuck, G.N. (ed.). A comprehensive study of the ecological impacts of the worldwide pelagic longline industry: southern hemisphere studies. CSIRO Marine Research, Hobart. Pp. 149-223.

Ryan, P.G. 2003. Estimating the demographic benefits of rehabilitating oiled African Penguins. In: Nel, D.C. and Whittington, P.A. (eds), Rehabilitation of oiled African Penguins: a conservation success story. BirdLife South Africa and Avian Demography Unit, Cape Town, pp. 25-29.

Glass, J.P. and Ryan, P.G. 2003. Conservation challenges in small communities: conservation management in the Tristan islands. In: Pienkowski, M. (ed.) A sense of direction: a conference on conservation in UK Overseas Territories and other small island communities. UK Overseas Territories Conservation Forum, www.ukotcf.org. Pp. 139-147.

Ryan, P.G. and Glass, J.P. 2001. Inaccessible Island Nature Reserve Management Plan. Government of Tristan da Cunha, Edinburgh, Tristan da Cunha. 65 pp.

Ryan, P.G., Purves, M. and Cooper, J. 2001. Problems with pirates: toothfish longlining and seabird bycatch at the Subantarctic Prince Edward Islands. In: Melvin, E.F. and Parrish, J.K. (eds) Seabird bycatch: trends, roadblocks, and solutions. University of Alaska Sea Grant, Fairbanks. Pp. 195.

Bloomer, P., Ryan, P.G. and Crowe, T.M. 1998. Some surprises from a molecular phylogeny for southern African larks. Ostrich 69: 397 [International Ornithological Congress abstract]

Nel, D.C., Hurford, J. and Ryan, P.G. 1998. Why is there sex-biased mortality of seabirds in the Prince Edward Island toothfish fishery? Ostrich 69: 442 [International Ornithological Congress abstract]

Ryan, P.G. and Marshall, H. 1998. The clapping arms race: why do some Clapper Larks not bother to play the game? Ostrich 69: 266 [International Ornithological Congress abstract]

Turpie, J.K. and Ryan, P.G. 1998. The value of birds and birding in South Africa. Ostrich 69: 455-456 [International Ornithological Congress abstract]

Ryan, P.G. 1997. Family Diomedeidae: albatrosses. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. and Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 13-15, 755-757.

Ryan, P.G. 1997. Family Procellariidae: petrels, shearwaters and prions. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. and Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 16-20, 757-761.

Ryan, P.G. 1997. Family Oceanitidae: storm petrels. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. and Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 21-23, 761-762.

Ryan, P.G. 1997. Family Phaethontidae: tropicbirds. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. and Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 762.

Ryan, P.G. 1997. Family Fregatidae: frigatebirds. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. and Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 763.

Ryan, P.G. 1997. Family Chionididae: sheathbills. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. and Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 772.

Ryan, P.G. and Crawford, R.J.M. 1997. Family Sulidae: gannets and boobies. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. and Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 28-29, 762-763.

Ryan, P.G., Crawford, R.J.M. and Whittington, P.A. 1997. Family Spheniscidae: penguins. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. and Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 4-5, 755.

Ryan, P.G., Williams, A.J., Crawford, R.J.M., Underhill, L.G., Brooke, R.K., Herremans, M. and Simmons, R.E. 1997. Family Laridae: skuas, gulls and terns. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. and Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 459-484, 772-775.

Dean, W.R.J., Allan, D.G., Herremans, M., Ryan, P.G. and Cohen, C. 1997. Family Alaudidae: larks. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. and Brown, C.J. (eds) The atlas of southern African birds. Vol. 2: Passerines. BirdLife South Africa, Johannesburg. Pp. 4-47, 687.

Gregory, M.R. and Ryan, P.G. 1997. Pelagic plastics and other seaborne persistent synthetic debris: a review of Southern Hemisphere perspectives. In: Coe, J.M. and Rogers, D.B. (eds) Marine debris: sources, impacts and solutions. Springer-Verlag, New York. Pp. 49-66.

Ryan, P.G. 1996. Identifying problem species: the Karoo Lark complex (Karoo, Red, Dune and Barlow’s Larks). In: Sinclair, J.C. and Hockey, P.A.R. The larger illustrated guide to birds of southern Africa. Struik, Cape Town. Pp. 423-424.

Allan, D.G. and Ryan, P.G. 1996. Morphometrics, sex ratio, molt, and stomach contents of Blue Cranes in the western Cape Province, South Africa. In: Beilfuss, R.D., Tarboten, W.R. and Sichuki, N.N. (eds) Proc. 1993 Afr. Crane and Wetland Training Workshop. International Crane Foundation, Baraboo, Wisconsin. Pp. 377-387.

Ryan, P.G., Hood, I., Bloomer, P. and Komen, J. 1995. Barlow’s Lark - a near endemic to the Sperrgebiet. Pp. 54-55 in Pallett, J. (ed.) The Sperrgebiet: Namibia’s least known wilderness. NAMDEB, Windhoek.

Cooper, J. and Ryan, P.G. 1995. The conservation status of Gough Island. In: Dingwall, P.R. (ed.) Progress in the conservation of the Subantarctic Islands. World Conservation Union: Conservation of the Southern Polar Regions 2: 71-84.

Cooper, J., Ryan, P.G. and Andrew, T.G. 1995. The conservation status of the Tristan da Cunha Islands. In: Dingwall, P.R. (ed.) Progress in the conservation of the Subantarctic Islands. World Conservation Union: Conservation of the Southern Polar Regions 2: 59-70.

Moloney, C.L and Ryan, P.G. 1995. Antarctic marine foodwebs. In: Nierenberg, W.A. (ed.) Encyclopedia of Environmental Biology, Vol. 1. Academic Press, San Diego. Pp 53-69.

Ryan, P.G. and Siegfried, W.R. 1994. The viability of small populations of birds: an empirical investigation of vulnerability. In: Remmert, H. (ed.) Minimum animal populations. Ecological Studies 106: 3-22. Springer-Verlag, Berlin. [reprinted in German and English in Nationalpark Berchtesgaden Forschungsbericht 27: 6-31.]

Cooper, J. and Ryan, P.G. 1994. Management Plan for the Gough Island Wildlife Reserve. Government of Tristan da Cunha, Edinburgh, Tristan da Cunha. 96 pp.

Crowe, T.M., Ryan, P.G., Essop, M.F., Brooke, R.K., Hockey, P.A.R. and Siegfried, W.R. 1994. Species as the 'currency' of conservation: the Karoo/Dune/Red Lark complex of south-western Africa. Pp 229-234 in Forey, P.L., Humphries, C.J. and Vane-Wright, R.I. (eds) Systematics and Conservation Evaluation. Clarendon Press, Oxford.

Ryan, P.G. 1990. The marine plastic debris problem off southern Africa: types of debris, their environmental effects, and control measures. Proc. Second Int. Conf. on Marine Debris, NOAA Tech. Mem., NMFS, SWFC 154: 85-102.

Ryan, P.G. 1990. The effects of ingested plastic and other marine debris on seabirds. Proc. Second Int. Conf. on Marine Debris, NOAA Tech. Mem., NMFS, SWFC 154: 623-634.

Ryan, P.G. and Rose, B. 1989. Migrant seabirds. In: Payne, A.I.L. and Crawford, R.J.M. (Eds). Oceans of life off southern Africa. Vlaeberg, Cape Town, pp. 274-287.

Popular articles (284):

Ryan, P. 2023. Adapt-a-bill. Yet more cryptic prions. African Birdlife 11(2): 10-11.

Ryan, P. 2023. Contact tracing. Feather lice transmission between seabirds. African Birdlife 11(2): 12.

Ryan, P. 2023. Change your tune. Coping with noisy neighbours. African Birdlife 11(2): 13.

Ryan, P. 2023. Cause and effect. Drop in Lesser Kestrel numbers. African Birdlife 11(2): 14.

Ryan, P. 2023. Príncipe Scops Owl. It’s official. African Birdlife 11(2): 15.

Ryan, P. 2023. Synthetic fibre blues. African Birdlife 11(2): 16.

Ryan, P. 2023. Canon R7 camera. Premium performance at a reasonable price. African Birdlife 11(2): 58-60.

Ryan, P. 2022. Mating moves: Meek male albatrosses don't get it. African Birdlife 11(1): 14.

Ryan, P.G. 2022. Owl origins: into the black (and back). African Birdlife 11(1): 15.

Ryan, P. 2022. Obituary: William Richard John Dean. African Birdlife 11(1): 18-19.

Ryan, P. 2022. Bird of the year 2022: Cape Gannet. Close quarters. African Birdlife 11(1): 37.

Ryan, P. 2022. All I want: Zeiss binoculars. African Birdlife 11(1): 60. 

Ryan, P.G. 2022. Bird of the year 2022: Cape Gannet. On the move. African Birdlife 10(6):50.

Ryan, P.G. 2022. Rapid eye movement: Field testing Canon’s R3 camera. African Birdlife 10(6):60-63.

Ryan, P.G. 2022. Desert enigma: Gray’s Lark. African Birdlife 10(6):12-13.

Ryan, P.G. 2022. Adapt and dive: penguin evolution and conservation. African Birdlife 10(6):10-11.

Ryan, P.G. and Marais, E. 2022. Mozambique: waiting to be explored. African Birdlife 10(6): 36-45.

Ryan, P.G. and Miller, S. 2022. Final review. African Birdlife 10(6): 16-17.

Ryan, P.G. 2022. Food on the fly: Red-footed Boobies in hot pursuit of flying fish. African Birdlife 10(5): 14-16. 

Pichegru, L. and Ryan, P.G. 2022. Huge volume of plastic flood debris trapped inshore by Agulhas Current. Daily Maverick: 14 June

Ryan, P.G. 2022. Bird of the year 2022: Cape Gannet. Investing in the future. African Birdlife 10(5): 60. 

Ryan, P.G. 2022. Conserving forest birds: The benefits of protected area status. African Birdlife 10(4): 14. 

Ryan, P.G. 2022. Too soon to panic: Are false alarm flights beneficial? African Birdlife 10(4): 15. 

Ryan, P.G. 2022. The FitzPatrick Report: Gough Island blues. African Birdlife 10(4): 18. 

Ryan, P.G. 2022. Ripple effect. African Birdlife 10(4): 26-27. 

Ryan, P.G. 2022. Bird of the year 2022: Cape Gannet. Team work. African Birdlife 10(4): 42. 

Ryan, P.G. 2022. On track: hidden diversity in Bar-tailed Godwits. v 10(3): 10. 

Ryan, P.G. and Peacock, F. 2022. Pale reflection: juvenile plumage of the Protea Canary.African Birdlife 10(3): 13. 

Ryan, P.G. 2022. The FitzPatrick Report: Ticking boxes. African Birdlife 10(3): 18. 

Ryan, P.G. 2022. Spectacular! The recovery of the Spectacled Petrel population. African Birdlife 10(3): 30–32. 

Ryan, P.G. 2022. Bird of the year 2022: Cape Gannet. Strike force. African Birdlife 10(3): 58-59. 

Ryan, P.G. 2022. Toxic overload: you are what you eat. African Birdlife 10(2): 12–13. 

Ryan, P.G. 2022. Urban perils: House Sparrows on the decline. African Birdlife 10(2): 10–11. 

Ryan, P.G. 2022. Tern about. African Birdlife 10(2): 16. 

Ryan, P.G. 2022. Sharp shooting: Canon’s RF 100–500 mm zoom lens. African Birdlife 10(2): 62–64. 

Ryan, P.G. 2022. Bird of the year 2022: Cape Gannet. Clear and present dangers. African Birdlife 10(2): 50–51. 

Ryan, P.G. and Vrettos, M. 2021. Stay or go: the evolutionary consequences of migration. African Birdlife 10(1): 24-29.

Ryan, P.G. 2021. Resetting the system. African Birdlife 10(1): 18–19. 

Ryan, P.G. 2021. On the shelf: Seabirds – the new identification guide. African Birdlife 10(1): 14–15. 

Ryan, P.G. 2021. The dark side: Do dark upperwings improve flight performance? African Birdlife 10(1): 12. 

Ryan, P.G. 2021. Birds to watch: Campbell Albatross off Africa. African Birdlife  9(6): 14.

Ryan, P.G. 2021. Sweet tooth. African Birdlife 9(6): 15.

Ryan, P.G. 2021. Feather light; leucistic White-fronted Plover – or is it? African Birdlife 9(5): 10-11.

Ryan, P.G. 2021. Birds to watch: American Cliff Swallow. African Birdlife 9(5): 13.

Ryan, P.G. 2021. Altogether now! Living in groups makes you clever. African Birdlife 9(5): 14.

Ryan, P.G. 2021. Cleaning house. African Birdlife 9(4): 18.

Ryan, P.G. 2021. Shore leave: beach closure allows Cape Cormorants to rest on Muizenberg Beach. African Birdlife 9(4): 13.

Ryan, P.G. 2021. True reflection: Canon’s R6 and R5 camera bodies. African Birdlife 9(3): 56-60.

Ryan, P.G. 2021. The ultimate point-and-shoot: Canon’s R6 and RF 800mm f11 IS STM lens. African Birdlife 9(3): 61.

Ryan, P.G. 2021. Sex and the single plover. African Birdlife 9(2): 24-27.

Ryan, P.G. 2021. Old baldy: Mite infestation in birds. African Birdlife 9(2): 15.

Ryan, P.G. 2021. Feeling chirpy. African Birdlife 9(2): 13.

Ryan, P.G. 2020. Windswept. African Birdlife 9(1): 12-13.

Ryan, P.G. 2020. Better and better: Zeiss SF 10x32 binoculars. African Birdlife 9(1): 60-61.

Ryan, P.G. 2020. On the flip side. African Birdlife 8(6): 10.

Ryan, P.G. 2020. Adapting to the new normal. African Birdlife 8(6): 12-13.

Ryan, P.G. 2020. Rate of return. African Birdlife 8(5): 12-14.

Ryan, P.G. 2020. Lessons in Lockdown: Covid-19 – a boon for birds? African Birdlife 8(5): 18-21.

Ryan, P.G. 2020. We looked for South Africa’s ‘missing’ plastic litter. This is what we found. https://theconversation.com/we-looked-for-south-africas-missing-plastic-litter-this-is-what-we-found-139782

Ryan, P.G. 2020. Puzzling prions. African Birdlife 8(4): 12-13.

Ryan, P.G. 2020. Supernormal clutches in Hartlaub’s Gulls. African Birdlife 8(4): 14.

Ryan, P.G. 2020. Shrinking returns: Long-term changes in northern Cape coastal birds. African Birdlife 8(4): 15.

Ryan, P.G. 2020. Vampire ravens. African Birdlife 8(3): 12.

Ryan, P.G. 2020. The more things change... African Birdlife 8(2): 12.

Ryan, P.G. 2020. Yet more greenbuls? African Birdlife 8(2): 13.

Ryan, P.G. 2020. Mix and match: hybrid Orange-breasted × Southern Double-collared Sunbird. African Birdlife 8(2): 14-15.

Ryan, P.G. 2020. Black out: the Canary Islands Oystercatcher re-examined. African Birdlife 8(2): 16.

Ryan, P.G. 2020. Message in a bottle. African Birdlife 8(2): 18-19.

Ryan, P.G. 2019. Bottleneck: by the skin of their non-existent teeth. African Birdlife 7(6): 10.

Ryan, P., Ludynia, K. and Pichegru, L. 2019. Ships’ risky fuel transfers are threatening African Penguins. https://theconversation.com/ships-risky-fuel-transfers-are-threatening-african-penguins-121575

Ryan, P.G. 2019. Annual reporting. African Birdlife 7(5): 16-17.

Ryan, P.G. 2019. Plastic poses a major environmental threat: but is it being over-stated? http://theconversation.com/plastic-poses-a-major-environmental-threat-but-is-it-being-over-stated-120175

Ryan, P.G. 2019. Testing, testing – too much knowledge can be a dangerous thing... African Birdlife 7(5): 20-21.

Ryan, P.G. 2019. Cotton on. African Birdlife 7(5): 38.

Ryan, P.G. 2019. Island survivor. African Birdlife 7(5): 50-55.

Ryan, P.G. 2019. Hand across the seas – another Africa-New Zealand link. African Birdlife 7(4): 11.

Ryan, P.G. 2019. Measuring research impact. African Birdlife 7(3): 18.

Ryan, P.G. 2019. Review: Leica APO Televid 82 with 25-50x WW eyepiece African Birdlife 7(3): 56-57.

Ryan, P.G. 2019. Expedition to Inaccessible Island. Tristan da Cunha Newsletter, February 2019: 26-29.

Ryan, P.G. 2019. Origins of the world’s smallest bird solved. Tristan da Cunha Newsletter, February 2019: 25

Ryan, P.G. 2019. Travelling trogons. African Birdlife 7(2): 10.

Ryan, P.G. 2019. Dicing with death. African Birdlife 7(2): 11.

Ryan, P.G. 2018. Going up... and down. African Birdlife 7(1): 10-11.

Ryan, P.G. 2018. Fact finding: the IOC and big data. African Birdlife 7(1): 18-20.

Ryan, P.G. 2018. Review: Kowa Genesis 33 Prominar 10x33 binoculars. African Birdlife 7(1): 73.

Ryan, P.G. 2018. What limits long-range migrants? African Birdlife 6(6): 10-11.

Ryan, P.G. 2018. The high cost of tracking. African Birdlife 6(6): 11.

Ryan, P.G. 2018. Home on loan. African Birdlife 6(6): 24-27.

Ryan, P.G. 2018. What a waste: plastic pollution update. African Birdlife 6(6): 32-41.

Ryan, P.G. 2018. Check your mate. African Birdlife 6(4): 10.

Ryan, P.G. 2018. Duck, down – White-backed Duck diving behaviour. African Birdlife 6(4): 12.

Ryan, P.G. 2018. Jouanin’s Petrel – or is it? African Birdlife 6(3): 10-11.

Risi, M., Jordaan, R. and Ryan, P. 2018. First record of Chilean Skua in the Indian Ocean. African Birdlife 6(3): 22.

Ryan, P.G. 2018. Review: Swarowski CL Companion binoculars. African Birdlife 6(3): 72-73.

Ryan, P.G. 2018. On the shelf: The Birder’s Guide to Africa. African Birdlife 6(2): 14-15.

Ryan, P.G. 2018. On the shelf: Shorebirds in Action. African Birdlife 6(2): 15.

Ryan, P.G. 2018. Review: Zeiss Conquest Gavia 85 T spotting scope. African Birdlife 6(2): 69.

Ryan, P.G. 2017. The FitzPatrick Report: Science and serendipity. African Birdlife 6(1): 18.

Ryan, P.G. 2017. Review: Warriers, Dilettantes & Businessmen. African Birdlife 6(1): 15.

Ryan, P.G. 2017. Short-tailed Shearwaters in the Atlantic Ocean. African Birdlife 5(6): 10-11.

Ryan, P.G. 2017. Binocular review – Leica Noctivid. African Birdlife 5(6): 74-75.

Ryan, P.G. 2017. Penguins through the ages. African Birdlife 5(5): 10.

Ryan, P.G. and Walton, J. 2017. Obituary – Rob Martin. African Birdlife 5(5): 40.

Ryan, P.G. 2017. To the edge and back – celebrating seabirds. African Birdlife 5(3): 52-58.

Ryan, P.G. 2017. Obituary – Barrie Rose 1947-2016. African Birdlife 5(3): 60.

Ryan, P.G. 2017. Surfers in action: inter-tidal clean-ups at Surfers’ Corner. Submerge (February-March 2017): 44.

Ryan, P.G. 2016. New scops owl on Príncipe. African Birdlife 5(1): 10.

Ryan, P.G. 2016. Sleeping with one eye open. African Birdlife 5(1): 22-25.

Ryan, P.G. 2016. Talking Tech: Crops and pixels. African Birdlife 5(1): 70-71.

Ryan, P.G. 2016. Flying colours. Wild Spring 2016: 68-73.

Ryan, P.G. 2016. Crash-test dummies: foraging ecology of Cape Gannets. African Birdlife 4(6): 22-30.

Ryan, P.G. 2016. See-saw: contrasting population trends in Phoebetria albatrosses. African Birdlife 4(5): 10-11.

Ryan, P.G. 2016. Many a bird’s rise from the ashes uncertain. SANParks Times (June 2016): 25.

Ryan, P.G. 2016. Review: Birds of Botswana. African Birdlife 4(4): 16.

Ryan, P.G. 2016. Another year, another tree. African Birdlife 4(4): 14.

Ryan, P. and Graham, J. 2016. Review: Nikon goes big. African Birdlife 4(3): 66-67.

Flood, R. and Ryan, P.G. 2016. Aging Shy Albatrosses. African Birdlife 4(2): 49-53.

Ryan, P.G. 2016. Pocket rockets: point and shoot cameras. African Birdlife 4(2): 44-47.

Ryan, P.G. 2016. World Seabird Conference making waves. African Birdlife 4(2): 13.

Ryan, P.G. 2015. Review: Canon’s new 36x12 IS III image-stabilised binoculars. African Birdlife 4(1): 68.

Ryan, P.G. 2015. Choosing your lens. African Birdlife 3(6): 44-52.

Ryan, P.G. 2015. Aliens and islands. Quest 11(3): 36-40.

Ryan, P.G. 2015. Gender bender: Sentinel Rock Thrush. African Birdlife 3(5): 14.

Barnard, P. and Ryan, P.G. 2015. Fire in the city. African Birdlife 3(4): 14-16.

Ryan, P.G. 2015. Birds on the Edge: the cost of conserving species. African Birdlife 3(3): 12.

Ryan, P.G. 2015. The last word on the evolution of birds. African Birdlife 3(3): 10-11.

Ryan, P.G. 2015. Man at work. African Birdlife 3(2): 44-51.

Ryan, P.G. 2015. Get your groove on. African Birdlife 3(2): 14.

Ryan, P.G. 2015. Practice makes perfect. African Birdlife 3(2): 12-13.

Ryan, P.G. 2014. Baby on board: Mascarene Petrel unveiled. African Birdlife 3(1): 10-11.

Ryan, P.G. 2014. Scaling… up and down. African Birdlife 3(1): 11.

Ryan, P.G. 2014. Follow the leader. Wild Spring 2014: 84-88.

Ryan, P.G. 2014. Calling time. African Birdlife 2(6): 11.

Ryan, P.G. 2014. Striking a balance: moulting flight feathers. African Birdlife 2(5): 56-60.

Ryan, P.G. 2014. Flying ostriches. African Birdlife 2(5): 18.

Ryan, P.G. 2014. Colour coding: Black power. African Birdlife 2(5): 14.

Ryan, P.G. 2014. Riders on the storm. African Birdlife 2(5): 10-11.

Ryan, P.G. 2014. Little Brown Jobs’ life on the edge. SANParks Times (June 2014): 26.

Ryan, P.G. 2014. Shed load: feather growth and replacement strategies. African Birdlife 2(4): 36-40.

Ryan, P.G. 2014. Orientation course: check the compass, chaps. African Birdlife 2(4): 12.

Ryan, P.G. 2014. A narrow margin: new prion discovered on Gough Island. African Birdlife 2(4): 10-11.

Ryan, P.G. 2014. Who’s a clever bird? Wild Autumn 2014: 86-89.

Cooper, J. and Ryan, P.G. 2014. Progress with supporting the Albatross and Petrel Agreement on the outer islands of Tristan da Cunha. Tristan da Cunha Newsletter 54: 32-34.

Ryan, P.G. 2014. Long distance flyers face tough times. SANParks Times (March): 22.

Ryan, P.G. 2014. Moult – the cost of annual renewal. African Birdlife 2(3): 37-43.

Ryan, P.G. 2014. …And cunning crocodiles. African Birdlife 2(3): 16.

Ryan, P.G. 2014. Do intelligent birds stress less? African Birdlife 2(3): 16.

Hockey, P.A.R. and Ryan, P.G. 2014. All together now: why do birds form groups? African Birdlife 2(2): 32-38.

Ryan, P.G. 2014. Nightjars in flux. African Birdlife 2(2): 13.

Ryan, P.G. 2013. Eagle eye. Wild Summer 2013/14: 72-75.

Ryan, P.G. 2013. Up, up and away. Wild Spring 2013:70-73.

Ryan, P. 2013. Mini marvels. African Birdlife 2(1): 30-31.

Ryan, P. 2013. The world’s most versatile wildlife lens? African Birdlife 2(1): 28-29.

Ryan, P. 2013. In the blink of an eye. African Birdlife 2(1): 11.

Ryan, P. 2013. Tackling mice on Gough. African Birdlife 2(1): 10-11.

Ryan, P.G. 2013. Party animals. Wild Winter 2013: 76-79.

Ryan, P. 2013. The eyes have it. African Birdlife 1(6): 28-34.

Ryan, P. 2013. Fear makes for faster babies. African Birdlife 1(6): 10.

Ryan, P. 2013. Underwater heron – African Darter. African Birdlife 1(5): 54-60.

Ryan, P. 2013. Tundra-breeding seabirds: the southern African connection. African Birdlife 1(5): 12.

Ryan, P. 2013. Tools of the trade: the new Canon 500-mm telephoto lens. African Birdlife 1(4): 22-23.

Ryan, P. 2013. There’s life in the old bird: geriatric sex and aging. African Birdlife 1(4): 12-13.

Ryan, P. 2013. Cinnamon-breasted Warbler’s Eastern Arc link. African Birdlife 1(4): 8.

Ryan, P. 2013. Creature comforts. Wild Autumn 2013: 90-92.

Ryan, P. 2013. Convergent evolution: binocular review 2013. African Birdlife 1(3): 48-52.

Ryan, P. 2013. Tool time: bait fishing by a Little Bittern. African Birdlife 1(3): 47.

Ryan, P. 2013. Parrots and falcons are sister to passerines. African Birdlife 1(3): 10.

Ryan, P. 2013. Swift moves. African Birdlife 1(2): 13.

Ryan, P. 2013. Splitting image: Cape, Kelp or Khoisan? African Birdlife 1(2): 10-11.

Ryan, P. 2013. Toxic waste: oceans of plastic threaten seabirds. African Birdlife 1(2): 52-56.

Ryan, P. 2012. Seabirding in the Straits of Malacca and Bay of Bengal. Suara Enggang 20(3): 6-7.

Ryan, P. 2012. African Penguins. African Birdlife 1(1): 30-31.

Ryan, P. 2012. When push comes to shove. African Birdlife 1(1): 6-8.

Ryan, P. 2012. Who’s that birder? Profiling avitourism in South Africa. Africa Birds and Birding 17(2): 52-53.

Melo, M. and Ryan, P. 2012. Endemism gone wild: the forgotten bird islands of São Tomé and Príncipe, Gulf of Guinea. Africa Birds and Birding 17(2): 32-41.

Ryan, P., Melo, M. and Stervander, M. 2012. Multinational scientific visit to Inaccessible Island. Tristan da Cunha Newsletter (February 2012): 32-33.

Ryan, P. 2012. Gujarat: A taste of Africa. Africa Birds and Birding 17(1): 58-65.

Ryan, P. 2012. Shoebill trade: the cost of being unique. Africa Birds and Birding 17(1): 15.

Ryan, P. 2012. Killer moves: save the albatross, club that seal. Africa Birds and Birding 17(1): 14-15.

Ryan, P. 2011. Bangweulu in the balance. Africa Birds and Birding 16(5): 66-67.

Ryan, P. 2011. User’s guide: Zeiss Victory PhotoScope 85 T*FL. Africa Birds and Birding 16(4): 16-17.

Ryan, P. 2011. Crisis in Utopia. Africa Birds and Birding 16(3): 16-17.

Ryan, P. 2011. Tracking the longest migration. Africa Birds and Birding 16(3): 15.

Ryan, P. 2011. Subantarctic Skuas in trouble? Africa Birds and Birding 16(3): 14.

Ryan, P. 2011. Slip sliding away - birds on the move. Africa Birds and Birding 16(1): 30-34.

Ryan, P. 2010. Best value-for-money binoculars. Africa Birds and Birding 15(6): 51-54.

Goren, M. and Ryan, P. 2010. Tristan Albatrosses off South Africa. Africa Birds and Birding 15(5): 14.

Cohen, C., Ryan, P.G., Claasen, M. and Ntoyinka, C. 2010. Birding Rwanda—in search of Red-collared Babbler Kupeornis rufocinctus, Albertine Rift endemics and more. Bulletin of the African Bird Club 17: 229-237.

Ryan, P. 2010. Tough times for seabirds. Africa Birds and Birding 15(4): 30.

Ryan, P. 2010. Resolving the enigmas: African oddities on the avian evolutionary tree. Africa Birds and Birding 15(3): 37-40.

Ryan, P. 2010. Dumped and bumped: chick recognition and rejection. Africa Birds and Birding 15(3): 14.

Ryan, P. 2010. Divide and rule: confusing times for seabirders. Africa Birds and Birding 15(3): 12.

Ryan, P. 2010. Tipping point: mice eroding Gough’s seabirds. Africa Birds and Birding 15(2): 13.

Ryan, P. 2010. Reproductive skew in African starlings. Africa Birds and Birding 15(2): 12.

Ryan, P. and Hockey, P. 2010. African Openbills on the move. Africa Birds and Birding 15(2): 8-9.

Ryan, P.G. and Cohen, C. 2010. Bridging the Rift: birding Rwanda. Africa Birds and Birding 15(1): 30-37.

Ryan, P.G. 2009. Sign of the times for Tristan Albatrosses. Africa Birds and Birding 14(3): 12.

Ryan, P.G. 2009. Tried and tested: Zeiss DC4 camera eyepiece. Africa Birds and Birding 14(2): 18-19.

Ryan, P.G. 2009. Sixth albatross breeding on Prince Edward Island. Africa Birds and Birding 14(2): 14.

Ryan, P.G. 2009. A strange twist for New Zealand’s Yellow-eyed Penguin. Africa Birds and Birding 14(2): 15.

Ryan, P.G. 2009. Home ground advantage: working your patch. Africa Birds and Birding 14(1): 16-18.

Pichegru, L. and Ryan, P.G. 2009. Teetering on the brink. Africa Birds and Birding 14(3): 42-53.

Ryan, P.G. 2008. Defrosting the Southern Ocean: climate change and the birds of Antarctica. Africa Geographic 16 (9): 54-62.

Ryan, P.G. 2008. Liuwa in the wet: birding western Zambia. Africa Birds and Birding 13(6): 44-53.

Ryan, P.G. 2008. Cold play. Africa: Birds and Birding 13(4): 54-56.

Ryan, P.G. 2008. Its not love, actually. Africa: Birds and Birding 13(4): 42-49.

Ryan, P.G. 2008. Girl power. Africa: Birds and Birding 13(4): 21.

Ryan, P.G. 2008. Dead sexy: Road mortality and necrophilia in Namib Larks. Africa: Birds and Birding 13(1): 15.

Ryan, P.G. 2008. User’s guide: Binoculars - mid-range marvels. Africa: Birds and Birding 13(1): 56-57.

Sinclair, I. and Ryan, P.G. 2008. Birding an ancient land: Socotra. Africa: Birds and Birding 13(1): 60-69.

Ryan, P.G. 2007. User’s guide: Best buy - spotting scopes. Africa: Birds and Birding 12(6): 55-59.

Ryan, P.G. 2007. Algal slime threatens Wrybill. Africa: Birds and Birding 12(4): 15.

Ryan, P.G. 2007. In the family way – Notiomystidae: another new family. Africa: Birds and Birding 12(4): 14.

Ryan, P.G. 2007. Under siege: west coast gannets. African Wildlife 61(3): 38-40.

Ryan, P.G. 2007. Mountains in the sky – La Reunion. Africa: Birds and Birding 12(2): 44-49.

Ryan, P.G. 2007. The north/south divide: penguin split adds another species to the African list. Africa: Birds and Birding 12(2): 14.

Ryan, P.G. 2007. Going, going, gannet… tough times for Benguela seabirds. Africa: Birds and Birding 12(1): 30-35.

Ryan, P.G. 2006. User’s guide: Best buy binoculars. Africa: Birds and Birding 11(6): 59-63.

Ryan, P.G. 2006. Digital SLR. Africa: Birds and Birding 11(5): 53-56.

Ryan, P.G. 2006. Digiscoping. Africa: Birds and Birding 11(4): 67-69.

Ryan, P.G. 2006. Naude’s Nek: on a high in the Drakensberg. Africa: Birds and Birding 11(3): 28-31.

Ryan, P.G. 2006. Sunbird conundrum. Africa: Birds and Birding 11(3): 12.

Ryan, P.G. 2006. Flights of fancy. Africa Geographic 14(4): 18-19.

Ryan, P.G. 2006. The long haul. Africa: Birds and Birding 11(2): 52-59.

Ryan, P.G., Graham, J. and Sutherland, A.C. 2006. Pelagic birding trips in the southern Mozambique Channel. Bull. Afr. Bird Club 13: 197-204.

Ryan, P.G. and Graham, J. 2006. Ntsikeni Nature Reserve – an overlooked IBA. Africa: Birds and Birding 11(1): 12-13.

Ryan, P.G. 2005. Toughing it out: birding Sierra Leone. Africa: Birds and Birding 10(6): 32-38.

Ryan, P.G. 2005. The long and the short of it: are flamingos modified grebes? Africa: Birds and Birding 10(6): 11.

Ryan, P.G. 2005. On a wing and a prayer: birds at risk. African Wildl. 59(4): 28-31.

Ryan, P.G. 2005. Ringeyes – the Inaccessible Petrel. Africa: Birds and Birding 10(5): 62-69.

Ryan, P.G. 2005. Why penguins no longer follow ostriches. Africa: Birds and Birding 10(4): 66-70.

Sinclair, I., Spottiswoode, C., Cohen, C., Mills, M, Cassidy, R. and Ryan, P. 2004. Birding western Angola. Bull. African Bird Club 11: 152-160.

Ryan, P.G. 2004. Current threats: conserving albatrosses. Africa: Birds and Birding 9(1): 38-45.

Ryan, P.G. 2003. Hurricane threatens Bermuda’s petrels. Africa: Birds and Birding 8(6): 15.

Sinclair, I. and Ryan, P.G. 2003. Cuanza Sul: the heart of Angola. Africa: Birds and Birding 8(3): 43-50.

Ryan, P.G. 2003. Tristan da Cunha: a bird haven in the South Atlantic. National Geographic Bird Watcher Jan-Feb: 8-10.

Ryan, P.G. 2002. Prince Edward Island. Africa Geographic 10(4): 76-81.

Ryan, P.G. 2002. More than an island fling? Attempted rapes by Wandering Albatrosses. Africa: Birds and Birding 7(2): 15.

Ryan, P.G. and Cassidy, R. 2002. The ghost in the mavunda: birding north-west Zambia. Africa: Birds and Birding 7(6): 37-41.

Ryan, P.G. 2002. Tailorbirds: rare beauties of the mountains in the mist. National Geographic Bird Watcher Jan-Feb: 4-7.

Ryan, P.G. 2001. New bird for Africa: Chatham Albatross. Africa: Birds and Birding 6(4): 18.

Ryan, P.G. and Sinclair, J.C. 2001. The great skua debate. Africa: Birds and Birding 6(4): 16-17.

Ryan, P.G. 2001. Albatrosses: ocean wanderers. Undercurrents 6(2): 4-5.

Ryan, P.G. 2001. SA Agulhas pelagic trip: 9-11 August. Sandwatch 13(2): 10-12.

Ryan, P.G. 2001. All hands on deck to save albatrosses and petrels. Africa: Birds and Birding 6(2): 22-23.

Ryan, P.G. and Spottiswoode, C.N. 2001. Tailorbirds ‘rediscovered’ in Mozambique. Africa: Birds and Birding 6(5): 18-19.

Ryan, P.G. 2001. All at sea. Africa: Birds and Birding 6(1): 13.

Ryan, P.G. 2000. Do Wandering Albatrosses really wander? Africa: Birds and Birding 5(6): 13.

Ryan, P.G. 2000. Update: longlining Africa: Birds and Birding 5(6): 17.

Ryan, P.G. 2000. The great plastic bag debate: Is a ban on plastic bags the answer? African Wildl. 54(6): 7-8.

Anon. (Ryan, P.G. and Cooper, J.) 2000. Light stick litter – an alternative solution. Fishing Ind. News Sthn Afr. 1(4): 25.

Ryan, P.G. 2000. Separating albatrosses: Tristan or Wandering? Africa: Birds and Birding 5(4): 35-39.

Ryan, P.G. 2000. Gull relationships all at sea. Africa: Birds and Birding 5(4): 20-21.

Ryan, P.G. 2000. Is banning plastic bags the answer? Africa: Environment and Wildlife 8(6): 11-12.

Ryan, P.G. 2000. Fighting procumbent pearlwort on Gough Island. Africa: Environment and Wildlife 8(5): 24-25.

Ryan, P.G. 1999. He who stoops, conquers: the fastest bird? Africa: Birds and Birding 4 (2): 17.

Ryan, P.G. 1999. Mission Mount Namuli: a trail of rediscovery. Africa: Environment and Wildlife 7(3): 60-68.

Ryan, P.G. 1999. Spectacled Petrel Procellaria conspicillata. World Birdwatch 21(1): 24-25.

Ryan, P.G. Spottiswoode, C., Parker, V., Graham, J., Cohen, C. and Bento, C. 1999. The birds of Namuli, northern Mozambique: retracing Vincent’s footsteps. Bull. Africa Bird Club 6: 138-143.

Ryan, P.G. and Bloomer, P. 1999. A plethora of longbilled larks. Bird Numbers 8: 20-21.

Parker, V. and Ryan, P.G. 1999. Namuli Apalis: an expedition to Mount Namuli, Mozambique. Endangered Wildlife 32: 8-11.

Turpie, J. and Ryan, P. 1999. What are birders worth? The value of birding in South Africa. Africa: Birds and Birding 4(1): 64-68.

Ryan, P.G. 1998. Summer birding: the south-western Cape. Africa: Birds and Birding 3(6): 54-59.

Ryan, P.G. 1998. Dwarf Raven or Somali Crow? Africa: Birds and Birding 3 (1): 13.

Ryan, P.G. 1998. Spectacled Petrel: A new species instantly endangered. Africa: Birds and Birding 3 (2): 13.

Ryan, P.G. 1998. We are hanging them out to die. American Bird Watcher 11(3): 1, 8-9.

Ryan, P.G., Barnes, K. and Sinclair, I. 1998. Tanzania’s Eastern Arc: endemic hotspot of eastern Africa. Africa: Birds and Birding 3(3): 46-52.

Ryan, P.G. and Hockey, P.A.R. 1998. Books for Africa: a guide to the field guides. Africa: Birds and Birding 3(3): 22-23.

Ryan, P.G. 1997. Lumping and splitting: the complexities of bird taxonomy. Africa: Birds and Birding 2(6): 64-67.

Ryan, P.G. 1997. Skuas: Pirates of the oceans. Africa: Birds and Birding 2(5): 57-63.

Ryan, P.G. 1997. The Wandering Albatross: Icon of the oceans. Africa: Birds and Birding 2 (1): 28-37.

Ryan, P.G. 1997. Sheathbills shipping in... Africa: Birds and Birding 2 (4): 14.

Ryan, P.G. 1997. Swamphens, Takahes and the evolution of flightlessness. Africa: Birds and Birding 2 (5): 10.

Barnes, K., Boix, C. and Ryan, P.G. 1997. Longlining - southern African seabirds on the hook again. Bull. Africa Bird Club 4(1): 9.

Ryan, P.G. 1996. Ethiopia: Birding the roof of Africa. Africa: Birds and Birding 1(5): 35-41.

Ryan, P.G. 1996. Barlow’s Lark; a new endemic lark for southern Africa. Africa: Birds and Birding 1(4): 65-70.

Ryan, P.G., Komen, J. and Moloney, C.L. 1996. The land birds of the Sperrgebiet. Lanioturdus 29: 8-25.

Ryan, P.G. 1996. Dinner jacket strategies: Why Jackass Penguins wear striped tuxedos. Africa: Birds and Birding 1(3): 44-50.

Ryan, P.G. 1996. New nightjar from Ethiopia. Africa: Environment and Wildlife 4 (1): 20.

Ryan, P.G. 1996. World Heritage status for Gough Island. Africa: Birds and Birding 1 (1): 8.

Ryan, P.G. 1996. Private initiatives help Whitewinged Flufftail. Africa: Environment and Wildlife 3 (6): 18.

Ryan, P.G. 1996. Birding in Mozambique: orioles and akalats. Africa: Environment and Wildlife 4(1): 34-36.

Ryan, P.G. 1996. Resurrecting Barlow’s Lark. Africa: Environment and Wildlife 3 (4): 10.

Ryan, P.G. 1995. Exploitation of tropical hardwood trees in Mozambique. African Wildlife 49 (6): 18-19.

Ryan, P.(G.) and Moloney, C.(L.) 1995. Third year of litter study. S. A. Commercial Marine 3(4): 18.

Ryan, P.G. and Swanepoel, D. 1995. Updating marine debris research in South Africa. Marine Debris Worldwide 1: 6-7.

Ryan, P.(G.) and Turpie, J. 1995. Putting a price tag on rarity. Birding in Southern Africa 47: 92-93.

Ryan, P.G. 1994. The great heard bird debate. Birding in Southern Africa 46: 69.

Ryan, P.G. 1990. Some enigmatic canaries: a case for shotgun birding. Birding in Southern Africa 42: 58.

Ryan, P.G. 1989. Common Nighthawk: a potential vagrant to southern Africa. Birding in Southern Africa 41: 80-81.

Ryan, P.G. and Graham, J.M.D. 1989. The trials and tribulations of the second Hudsonian Godwit for southern Africa. Birding in Southern Africa 41: 103-104.

Ryan, P.G. and Watkins, B.P. 1989. First South African biological survey in Dronning Maud Land, Antarctica. BIOTAS Newsletter 4: 39.

Ryan, P.G. 1988. Plastic pollution: a serious threat to aquatic life. African Wildlife 42: 199-205.

Ryan, P.G. 1988. In search of the Inaccessible Island Rail and other insular pursuits. Excellence 4 (3): 61-64.

Ryan, P.G. 1987. Purple chickens and other tropical delights. Bokmakierie 39: 94-95.

Watkins, B.P. and Ryan, P.G. 1987. Mites, mosses and mumiyo. University of Cape Town Institute of Oceanography yearbook. 1987: 12-15.

Selected notes (33):

Ryan, P.G. and Walton, J. 2017. Obituary – Rob Martin. African Birdlife 5(5): 40.

Ryan, P.G. 2017. Obituary – Barrie Rose 1947-2016. African Birdlife 5(3): 60.

Ryan, P.G. 2015. First record of Lesser Yellowlegs Tringa flavipes for Tristan da Cunha, South Atlantic. Bulletin of the African Bird Club 22: 84-85.

Ryan, P.G. and Dyer, B.M. 2014. Kelp Gulls breeding in mountain fynbos. Ornithological Observations 5: 128-130.

Ryan, P.G. 2013. Obituary: Phil Hockey (1956-2013). Ibis 155: 698-700.

Ryan, P.G., Colville, J.F. and Picker, M. 2013. African Pipit eating monkey beetles. Ornithological Observations 4: 6-8.

Little R.M. and Ryan, P.G. 2013. Obituary: Philip Anthony Richard Hockey – ‘Education is what is left when what has been learnt has been forgotten’ (8 March 1956 – 24 January 2013). Ostrich 84: v.

Little R.M. and Ryan, P.G. 2013. Obituary: Phil Hockey: 8 March 1956 – 24 January 2013. South African Journal of Wildlife Research 43: 87.

Ryan, P.G. 2012. Blue-breasted Kingfisher Halcyon malimbica feeding at sea. Malimbus 34: 43-45.

Ryan, P.G., Shaw, J.M., van der Merwe, R. and van der Merwe, E. 2012. Pale Chanting Goshawks and other birds feeding on carrion. Ornithological Observations 3: 102-106.

Ryan, P.G. 2010. First record of Chilean Skua Catharacta chilensis for the African region Bulletin of the African Bird Club 17: 217-219.

Visser, P., Louw, H., Cuthbert, R. and Ryan, P. 2009. Salvin’s Albatross Thalassarche salvini on Gough Island, South Atlantic. Bull. Afr. Bird Club 16: 215-216.

Cooper, J., Ryan, P. and Glass, J. 2006. Eradicating invasive species in the United Kingdom Overseas Territory of Tristan da Cunha. Aliens 23: 1-3.

Ryan, P.G. 2002. Chatham Albatross Thalassarche eremita: new to Africa. Bull. African Bird Club 9: 44.

Ryan, P.G. and Sinclair, J.C. 1998. Somali Starlings Onychognathus blythii in south-central Ethiopia. Bull. Africa Bird Club 5: 56-57.

Wainger, L., Ryan, P., Cowling, R. and Costanza, R. 1996. Ecological economics in South Africa: the valuation and management of fynbos ecosystems. Ecol. Econ. Bull. 1(1): 12-13, 18, 20.

Cooper, J. and Ryan, P.G. 1995. SA scientists produce management plan for Gough Island. S. Afr. J. Sci. 91: 107.

Ryan, P.G. and Moloney, C.L. 1994. Range extension and display flight of Cuckoo Hawk. Gabar 9(2): 25.

Smith, P.W., Smith, S.A., Ryan, P.G. and Cassidy, R. 1994. First report of Virginia’s Warbler from the Bahama Islands, with comments on other records from the West Indies and eastern North America. El Pitirre 7 (2): 2-3.

Ryan, P.G. 1992. Fiscal Shrike feeding by plunge-diving. Ostrich 63: 42.

Ryan, P.G. and Moloney, C.L. 1991. Tristan Thrushes kill adult White-bellied Storm-Petrels. Wilson Bull. 103: 130-132.

Ryan, P.G. and Avery, G. 1990. Arctic Skua killed by sea-barbel spine. Br. Birds 83: 282-283.

Ryan, P.G. 1989. The distribution and abundance of Longtailed Skuas off southern Africa. Ostrich 60: 89-90.

Ryan, P.G. 1989. Common Nighthawk Chordeiles minor and new records of seabirds from Tristan da Cunha and Gough Islands. Bull. Br. Orn. Club 109: 147-149.

Ryan, P.G. 1988. Commentary: Concern about plastic pollution in Southern Ocean seabirds. Cormorant 16: 1-2.

Ryan, P.G., Peddemors, V.M. and Cockcroft, V.G. 1987. Dark morph Fregetta stormpetrel south of Africa. Cormorant 15: 91-92.

Ryan, P.G. 1986. An ocean of plastic. Conserva 1(3): 6-7.

Ryan, P.G. 1986. Records of skuas and Sabine’s Gulls ashore in southern Africa. Cormorant 13: 107-111.

Ryan, P.G. 1985. Antarctic Terns Sterna vittata feeding on the ground at Tristan da Cunha. Cormorant 13: 64-66.

Ryan, P.G. and Rose, B. 1985. Bradfield’s Swifts using cliffs and palm trees in Namibia. Ostrich 56: 218.

Ryan, P.G., Cassidy, R. and Salinger, T. 1984. The Haroni-Lusitu Junction revisited. Honeyguide 30: 90-91.

Ryan, P.G., Cassidy, R. and Salinger, T. 1984. Black and White Flycatcher: third record for Zimbabwe. Honeyguide 30: 127-128.

Ryan, P.G. and Graham, J.M.D. 1984. First record of the Lesser Yellowlegs in South Africa. Ostrich 55: 222-223.

Selected reports (22):

Ryan, P.G., Broome, K., Connan, M., de Bruyn, P.J.N., Fisher, P., Makhado, A., Oppel, S., Phillips, R.A. and Samaniego, A., with Wolfaardt, A. and Springer, K. 2023. An assessment of the risks of primary or secondary poisoning to non-target species that may result from the planned baiting operation at Marion Island, and advice on potential approaches to reduce these risks. Mouse-Free Marion, Johannesburg. 43 pp.

Ryan, P.G. 2018. Plastics and AEWA waterbirds: incidence and impacts. Unpublished report to the Agreement on the Conservation of African-Eurasian Migratory Waterbirds, 35 pp.

UNEP and WIOMSA 2008. Marine Litter in the Eastern Africa Region: An Overview Assessment. United Nations Environment Programme (Ryan, P.G. contributed the report for South Africa). Nairobi: UNEP Regional Seas Programme, 2008. 60 pp.

Ryan, P.G. 2007. Managing alien plants on the outer islands of Tristan da Cunha: follow-up flax eradication efforts. Report on activities: October-November 2007. Unpubl. rpt to UK Overseas Territory Environment Programme.

Ryan, P.G. 2007. Species status of the Spectacled Petrel Procellaria conspicillata. Unpubl. report to the Taxonomy Working Group of ACAP, Hobart, Tasmania.

Ryan, P.G. 2007. Yellow-nosed Albatross: one species or two? Unpubl. report to the Taxonomy Working Group of ACAP, Hobart, Tasmania.

Ryan, P.G. 2006. The current status of marine litter/debris in South Africa. Unpubl. rpt to the UN Environment Programme (WIOMSA). 43 pp.

Ryan, P.G., Barendse, J., Chiloane, L.A. and Moreku, G.L. 2004. Clearing invasive flax Phormium tenax on Inaccessible Island. Royal Society for the Protection of Birds, UK.

Cooper, J. and Ryan, P.G. 2004. South African National Plan of Action for reducing the incidental catch of seabirds in longline fisheries. Marine and Coastal Management, Department of Environmental Affairs and Tourism, Cape Town.

Cooper, J., Crawford, R.J.M., Nel, D.C., Petersen, S.L. and Ryan, P.G. 2004. Status and conservation of albatrosses and the larger petrels in South Africa. ACAP 1^st Meeting of Parties, ACAP/MOP1/IP.x

Ryan, P.G. and Watkins, B.P. 2001. Seabird bycatch in the Patagonian Toothfish longline fishery at the Prince Edward Islands: 2000-2001. CCAMLR-WG-FSA 01/XX.

Ryan, P.G. and Watkins, B.P. 2000. Seabird bycatch in the Patagonian Toothfish longline fishery at the Prince Edward Islands: 1999/2000. CCAMLR-WG-FSA 00/30.

Whittington, P.A., Crawford, R.J.M., Huyser, O., Oschadleus, D., Randall, R., Ryan, P., Shannon, L.V., Woolfardt, A., Cooper, J., Lacy, R.C. and Ellis, S. 2000. African Penguin population and habitat viability assessment. Final report. IUCN/SSC Conservation Breeding Specialist Group, Apple Valley MN (134 pp.).

Ryan, P.G. and Watkins, B.P. 1999. Seabird bycatch in the Patagonian Toothfish longline fishery at the Prince Edward Islands: 1998-1999. CCAMLR-WG-FSA 99/22.

Ryan, P.G. and Purves, M. 1998. Seabird bycatch in the Patagonian Toothfish longline fishery at the Prince Edward Islands: 1997-1998. CCAMLR-WG-FSA 98/36.

Ryan, P.G., Shannon, L.J., Crawford, R.J.M., Dyer, B.M. and Whittington, P.A. 1998. African Penguins at Boulders Coastal Park, Simon’s Town: the scientific basis for management. Percy FitzPatrick Institute unpubl. rpt for South African National Parks (72 pp.).

Turpie, J.K. and Ryan, P.G. 1998. The nature and value of birding in South Africa. BirdLife South Africa Research Report 1 (41 pp.).

Ryan, P.G., Boix-Hinzen, C., Enticott, J.W., Nel, D.C., Wanless, R. and Purves, M. 1997. Seabird mortality in the longline fishery for Patagonian Toothfish at the Prince Edward Islands: 1996-1997. CCAMLR-WG-FSA 97/51.

Ryan, P.G. 1995. The possible role of birds in causing transient earth faults on the Harvard-Merapi transmission line. Percy FitzPatrick Institute unpubl. rpt for ESKOM (23 pp.).

Ryan, P.G. and Moloney, C.L. 1995. Survey of marine debris stranded on South African beaches: winter 1994. Percy FitzPatrick Institute unpubl. rpt for Plastics Federation of South Africa (17 pp.).

Ryan, P.G. 1991. Comparison of the bird populations at three sites on the northwestern Cape coast. Percy FitzPatrick Institute unpubl. rpt for ESKOM (34 pp.).

Ryan, P.G. and Watkins, B.P. 1988. Biological survey of Robertskollen, western Dronning Maud Land, Antarctica. Percy FitzPatrick Institute unpubl. rpt for SASCAR (80 pp.).